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would have resulted in much natural mortality, limiting
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tree densities.
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Because of fire suppression, some pine rockland
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fragments now have a canopy of hardwoods. The most
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common canopy hardwood is wild-tamarind. Exotic
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hardwoods may also be common canopy trees in firesuppressed sites, including Australian umbrella tree and
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woman’s tongue .
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3.4.2 Subcanopy
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Subcanopy densities are now much denser and taller than they were under historical conditions.
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Conditions are site specific and vary according to geographic location, fire frequency, and
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distance to seed sources (especially rockland hammocks). Sites that have received frequent fires,
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such as Ludlam Pineland, are probably close to historic conditions, with short palm and
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Table 1: Seedling pine plantings
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on Miami-Dade County preserves
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Preserve Owner
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A.D. Barnes Park Parks
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Andrew Dodge Pineland DERM
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Black Creek Forest EEL
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Boystown Pineland EEL
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Deering Estate at Cutler Parks
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Eachus Pineland EEL
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Goulds Pineland EEL
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Larry & Penny Thompson Park Parks
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Ludlum Pineland EEL
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Martinez Pineland Parks
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Miami Metrozoo Parks
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Navy Wells Pineland Parks
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Ned Glenn Nature Preserve EEL
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Nixon Smiley Addition EEL
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Nixon Smiley Pineland Parks
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Palm Drive Pineland EEL
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Rockdale Pineland EEL
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Ron Ehman Park Parks
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Seminole Wayside Park Parks
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Trinity Pineland EEL
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Tropical Park Parks
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West Biscayne Pineland EEL
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EEL Program, Management Plan, Part II – Pine Rockland (DRAFT) _____________________________________________________________________________________________________________________
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Page 16
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hardwoods at low stem densities. Other sites, such as Navy Wells #23, are probably very
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different than they were historically. This site now has a tall understory of hardwoods with a
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very high stem density.
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3.4.3 Herb Layer
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The herb layer naturally has the highest diversity of plant species and a diverse array of rare
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plant species. Like the subcanopy layer, the herb layer varies greatly between forest fragments.
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Some sites that burned frequently and have sparse subcanopy layers have diverse herb layers.
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Other sites with dense subcanopies that result in low light penetration and thick duff
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accumulations have lost almost all herbaceous species. The overall herb layer in pine rockland
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fragments has been degraded, with a loss in density and diversity.
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3.4.4 Edges
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Historically, edges in the pine rockland ecosystem were limited to ecotones with other habitat
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types, especially rockland hammocks and marl prairies (see Sections 2.5 above and 3.5 below).
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Because of forest fragmentation, all pine rockland fragments now have edges with artificial
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communities, including lawns, agricultural fields, roads, and other vacant land. Edges create a
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unique ecotone that can have impacts to the interior of natural sites (See Part I). Edges of pine
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rockland fragments vary greatly in vegetation structure and composition, depending upon fire
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history, soil disturbance, and management history. These edges typically have dense growths of
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exotic and ruderal plant species. Abundant exotic plants include Brazilian pepper and Burma
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reed. Initial establishment of these exotic plants on edges allows their population sizes to build in
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disturbed soils, and then intact pine rocklands in the interior of the sites are subsequently invaded
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due to a heavy seed rain.
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3.5 Association with Other Habitat Types
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As discussed in Section 2.5, pine rockland was historically associated with other habitat types,
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especially rockland hammock and marl prairie. Today, most preserves have only a single habitat
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type and have no natural ecotones with other habitats. Exceptions on EEL sites include Silver
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Palm Groves, which has a small rockland hammock, and Nixon Smiley Pineland Addition,
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which has several small marl prairies. The loss of ecotone habitat has resulted in the loss of many
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populations of plant species restricted to the habitat, and has probably reduced the value of pine
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rockland for some wildlife species.
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3.6 Successional Processes Under Current Conditions
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Because of forest fragmentation, human intervention, and a mosaic of urban lands between pine
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rockland fragments, successional processes now differ than those discussed in Section 2.6. These
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changes are discussed below.
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3.6.1 Fire
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Since pine rocklands are fire climax communities, natural fire frequency is of great importance.
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However, fire periodicity, behavior, and intensity have all been changed on pine rockland
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fragments. Under historical conditions, pine rockland fragments burned every three (3) to seven
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(7) years, often in large landscape-scale, wind-driven fires. These fires were usually in the spring
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EEL Program, Management Plan, Part II – Pine Rockland (DRAFT) _____________________________________________________________________________________________________________________
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Page 17
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or summer. Following fragmentation, fire periodicity has lengthened considerably at most sites.
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Fire has been completely eliminated at some sites. This change can be attributed to two causes:
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intentional fire suppression by people and the elimination of large scale landscape-level fires
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because of the destruction of natural areas.
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In addition to periodicity, fire behavior has changed. Natural fires were usually wind-driven head
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fires that were ignited by lightning. Fires now start for a variety of reasons, including lightning,
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arson, accident, and by prescription. Many fires, especially prescribed fires, are backing fires
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burning into the wind. These are slow moving fires and can damage feeder roots of many plants.
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They are typically used because they are easier to control.
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Fire intensity has changed with the reduction in fire periodicity. Heavier fuel buildups between
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fires result in hotter temperatures and higher flames. Intense fires can have long-lasting impacts
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when they occur. Fires that are too intense can result in massive mortality of pines, saw
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palmettos, understory hardwoods, and herbs. Following the fires, sites can be invaded more
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readily by exotic pest plant species, especially natal grass, and ruderal species. Long term
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vegetation recovery following intense fires has not been studied.
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3.6.2 Tropical Cyclones
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There has been much recent discussion about global warming and hurricane activity. Some argue
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that global warming is resulting in stronger, more intense tropical cyclones (Webster et al. 2005).
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Some, however, argue against this (Hoyos et al. 2006). Regardless of this debate, the widespread
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changes in the overall landscape of Miami-Dade County and the changes to pine rockland
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