One of the later ruptures between Toghrul and Temüjin was Toghrul's refusal to give his daughter in marriage to Jochi, the eldest son of Temüjin, a sign of disrespect in the Mongolian culture. This act led to the split between both factions and was a prelude to war. Toghrul allied himself with Jamukha, who already opposed Temüjin's forces; however, the internal dispute between Toghrul and Jamukha, plus the desertion of a number of their allies to Temüjin, led to Toghrul's defeat. Jamukha escaped during the conflict. This defeat was a catalyst for the fall and eventual dissolution of the Keraite tribe.