Insects were among the earliest terrestrial herbivores and acted as major selection agents on plants. Plants evolved chemical defenses against this herbivory and the insects, in turn, evolved mechanisms to deal with plant toxins. Many insects make use of these toxins to protect themselves from their predators. Such insects often advertise their toxicity using warning colors. This successful evolutionary pattern has also been used by mimics. Over time, this has led to complex groups of coevolved species. Conversely, some interactions between plants and insects, like pollination, are beneficial to both organisms. Coevolution has led to the development of very specific mutualisms in such systems.