Many of the leading universities associated with Enlightenment progressive principles were located in northern Europe, with the most renowned being the universities of Leiden, Göttingen, Halle, Montpellier, Uppsala and Edinburgh. These universities, especially Edinburgh, produced professors whose ideas had a significant impact on Britain's North American colonies and, later, the American Republic. Within the natural sciences, Edinburgh's medical also led the way in chemistry, anatomy and pharmacology. In other parts of Europe, the universities and schools of France and most of Europe were bastions of traditionalism and were not hospitable to the Enlightenment. In France, the major exception was the medical university at Montpellier.