In 885 AD the Armenians reestablished themselves as a sovereign kingdom under the leadership of Ashot I of the Bagratid Dynasty. A considerable portion of the Armenian nobility and peasantry fled the Byzantine occupation of Bagratid Armenia in 1045, and the subsequent invasion of the region by Seljuk Turks in 1064. They settled in large numbers in Cilicia, an Anatolian region where Armenians were already established as a minority since Roman times. In 1080, they founded an independent Armenian Principality then Kingdom of Cilicia, which became the focus of Armenian nationalism. The Armenians developed close social, cultural, military, and religious ties with nearby Crusader States, but eventually succumbed to Mamluk invasions. In the next few centuries, Djenghis Khan, Timurids, and the tribal Turkic federations of the Ak Koyunlu and the Kara Koyunlu ruled over the Armenians.