London's defences were rapidly reorganised by General Pile, the Commander-in-Chief of Anti-Aircraft Command. The difference this made to the effectiveness of air defences is questionable. The British were still one-third below the establishment of heavy anti-aircraft artillery AAA (or ack-ack) in May 1941, with only 2,631 weapons available. Dowding had to rely on night fighters. From 1940 to 1941, the most successful night-fighter was the Boulton Paul Defiant; its four squadrons shot down more enemy aircraft than any other type. AA defences improved by better use of radar and searchlights. Over several months, the 20,000 shells spent per raider shot down in September 1940, was reduced to 4,087 in January 1941 and to 2,963 shells in February 1941.