Insects are the only invertebrates to have developed active flight capability, and this has played an important role in their success.:186 Their muscles are able to contract multiple times for each single nerve impulse, allowing the wings to beat faster than would ordinarily be possible. Having their muscles attached to their exoskeletons is more efficient and allows more muscle connections; crustaceans also use the same method, though all spiders use hydraulic pressure to extend their legs, a system inherited from their pre-arthropod ancestors. Unlike insects, though, most aquatic crustaceans are biomineralized with calcium carbonate extracted from the water.