In 2008, biochemist Boris Arkadievich Malyarchuk (Russian: Борис Аркадьевич Малярчук) et al. of the Institute of Biological Problems of the North, Russian Academy of Sciences, Magadan, Russia, used a sample (n=279) of Czech individuals to determine the frequency of "Mongoloid" "mtDNA lineages". Malyarchuk found Czech mtDNA lineages were typical of "Slavic populations" with "1.8%" Mongoloid mtDNA lineage. Malyarchuk added that "Slavic populations" "almost always" contain Mongoloid mtDNA lineage. Malyarchuk said the Mongoloid component of Slavic people was partially added before the split of "Balto-Slavics" in 2,000–3,000 BC with additional Mongoloid mixture occurring among Slavics in the last 4,000 years. Malyarchuk said the "Russian population" was developed by the "assimilation of the indigenous pre-Slavic population of Eastern Europe by true Slavs" with additional "assimilation of Finno-Ugric populations" and "long-lasting" interactions with the populations of "Siberia" and "Central Asia". Malyarchuk said that other Slavs "Mongoloid component" was increased during the waves of migration from "steppe populations (Huns, Avars, Bulgars and Mongols)", especially the decay of the "Avar Khaganate".