The study of altruism was the initial impetus behind George R. Price's development of the Price equation, a mathematical equation used to study genetic evolution.
ðə stˈʌdi ʌv ˈɔːltɹuːˌɪzəm wʌzðɪ ɪnˈɪʃəl ˈɪmpəɾəs bᵻhˌaɪnd dʒˈɔːɹdʒ ˈɑːɹ. pɹˈaɪsᵻz dɪvˈɛləpmənt ʌvðə pɹˈaɪs ᵻkwˈeɪʒən, ɐ mˌæθɪmˈæɾɪkəl ᵻkwˈeɪʒən jˈuːzd tə stˈʌdi dʒɪnˈɛɾɪk ˌɛvəlˈuːʃən.