The territory is marked by both sexes, as they both have developed anal glands from which they extrude a black substance that is smeared on rocks or grass stalks in -long streaks.
ðə tˈɛɹɪtˌoːɹi ɪz mˈɑːɹkt baɪ bˈoʊθ sˈɛksᵻz, æz ðeɪ bˈoʊθ hæv dɪvˈɛləpt ˈeɪnəl ɡlˈændz fɹʌm wˈɪtʃ ðeɪ ɛkstɹˈuːd ɐ blˈæk sˈʌbstəns ðæt ɪz smˈɪɹd ˌɔn ɹˈɑːks ɔːɹ ɡɹˈæs stˈɔːks ɪn lˈɔŋ stɹˈiːks.