The front feet have lost the pollex (or 'thumb'), resulting in four toes, while the rear feet have all five toes.
ðə fɹˈʌnt fˈiːt hæv lˈɔst ðə pˈɑːlɛks ɔːɹ θˈʌm, ɹɪzˈʌltɪŋ ɪn fˈoːɹ tˈoʊz, wˌaɪl ðə ɹˈɪɹ fˈiːt hæv ˈɔːl fˈaɪv tˈoʊz.