The salamanders left odour marks around their territories which averaged  in size and were sometimes inhabited by a male and female pair.
ðə sˈælɐmˌændɚz lˈɛft ˈoʊdɚ mˈɑːɹks ɚɹˈaʊnd ðɛɹ tˈɛɹɪtˌoːɹiz wˌɪtʃ ˈævɹɪdʒd ɪn sˈaɪz ænd wɜː sˈʌmtaɪmz ɪnhˈæbɪɾᵻd baɪ ɐ mˈeɪl ænd fˈiːmeɪl pˈɛɹ.