ɪn hɪz ɹᵻsˈɜːtʃ, hiː ˈaʊtlaɪnd ðə sɪɡnˈɪfɪkəns ʌvðə sˈɛntɹəl nˈɜːvəs sˈɪstəm ˌɔn mˈuːvmənt, dɹˈuː dɪstˈɪŋkʃənz bᵻtwˌiːn vˈɑːləntɚɹi ænd ɪnvˈɑːləntɚɹi ˈækʃənz ænd klˈæɹᵻfˌaɪd ðə kəmpˈoʊnənts ʌvðə lˈaɪt ɹˈiːflɛks wɪðˌɪn ðɪ ˈaɪ.
In his research, he outlined the significance of the central nervous system on movement, drew distinctions between voluntary and involuntary actions and clarified the components of the light reflex within the eye.