hiː wʌz ɐwˈɔːɹdᵻd ɐ pˌiːˈeɪtʃ dˈiː fɹʌmðə jˌuːnɪvˈɜːsᵻɾi ʌv kˈeɪmbɹɪdʒ ɪn nˈaɪntiːnhˈʌndɹɪd sˈɛvənti tˈuː fɔːɹ ɹᵻsˈɜːtʃ ɔnðə dʒɪnˈɛɾɪks ænd fˈʌŋkʃən ʌvðə meɪˈɑːɾɪk pˈɛɹɪŋkəntɹˈoʊl sˈɪstəmz sˈuːpɚvˌaɪzd baɪ ɹˈælf ɹˈaɪli.
He was awarded a PhD from the University of Cambridge in 1972 for research on the genetics and function of the meiotic pairing-control systems supervised by Ralph Riley.