hiː wʌz ɐsˈɪstənt, ɐsˈoʊsɪˌeɪt ænd fˈʊl pɹəfˈɛsɚɹ æt hˈɑːɹvɚd jˌuːnɪvˈɜːsᵻɾi ɪnðə dᵻpˈɑːɹtmənt ʌv sˈɛljʊlɚ ænd dɪvˈɛləpmˈɛntəl baɪˈɑːlədʒi fɹʌm nˈaɪntiːnhˈʌndɹɪd ˈeɪɾi fˈoːɹ nˈaɪntiːnhˈʌndɹɪd nˈaɪnti θɹˈiː ænd mˈuːvd tʊ ˈʌksd ænd ˌeɪtʃˌeɪtʃˌɛmˈaɪ ɪn nˈaɪntiːnhˈʌndɹɪd nˈaɪnti θɹˈiː.
He was Assistant, Associate and Full Professor at Harvard University in the Department of Cellular and Developmental Biology from 1984–1993 and moved to UCSD and HHMI in 1993.