ɪn hɪz ˈɜːli kɚɹˈɪɹ, hiː pˈʌblɪʃt ˌɔn dˈeɪnɪʃ ænd ˈɑːɹktɪk nˈɛmɐtˌoʊdz ænd ɔnðə zənˈeɪʃən ʌv mɚɹˈiːn ˈælɡiː ɪn ˈoːɹsʌnd.
In his early career, he published on Danish and Arctic nematodes and on the zonation of marine algae in Øresund.