ɪn nˈaɪntiːnhˈʌndɹɪd nˈaɪnti fˈaɪv hiː bɪkˌeɪm ðə fˈɪfθ sˌɜː wˈɪljəm dˈʌn pɹəfˈɛsɚɹ ʌv bˌaɪoʊkˈɛmɪstɹi ænd hˈɛd ʌvðə dᵻpˈɑːɹtmənt ʌv bˌaɪoʊkˈɛmɪstɹi æt ðə jˌuːnɪvˈɜːsᵻɾi ʌv kˈeɪmbɹɪdʒ; hiː kˈɜːɹəntli ˈɔːlsoʊ hˈoʊldz ðə tʃˈɛɹ ʌvðə skˈuːl ʌv bˌaɪəlˈɑːdʒɪkəl sˈaɪənsᵻz æt ðæt jˌuːnɪvˈɜːsᵻɾi.
In 1995 he became the fifth Sir William Dunn Professor of Biochemistry and head of the Department of Biochemistry at the University of Cambridge; he currently also holds the Chair of the School of Biological Sciences at that university.