hiː ðˈɛn mˈuːvd tə ðə jˌuːnɪvˈɜːsᵻɾi ʌv kˌɑːlɚɹˈɑːdoʊ bˈoʊldɚ fɔːɹ hɪz pˌiːˈeɪtʃ.dˈiː., wˌɛɹ hiː fˈɜːst dɪvˈɛləpt ðə dʒˌiːjˌuːˈɛs ɹᵻpˈoːɹɾɚ sˈɪstəm, ˈaɪsəlˌeɪɾɪŋ, sˈiːkwənsɪŋ ænd kˈæɹɪktɚɹˌaɪzɪŋ ðə fˈɜːst maɪkɹˈoʊbɪəl ɡlˈuːkjʊɹɹˌɑːnɪdˌeɪs, ænd kɹiːˈeɪɾɪŋ tɹænsdʒˈɛnɪk tɛknˈɑːlədʒi fɔːɹ kˌiːnoːɹhɐbdˈaɪɾɪs ˈɛlɪɡənz
He then moved to the University of Colorado Boulder for his Ph.D., where he first developed the GUS reporter system, isolating, sequencing and characterizing the first microbial glucuronidase, and creating transgenic technology for Caenorhabditis elegans