æt ðə sˈeɪm tˈaɪm ðə tʃˈɑːɹdʒᵻz ɔnðə dʒˌiːˌeɪdʒˈiː tʃˈeɪnz ɐtɹˈækt ˈaɪɑːnz ænd wˈɔːɾɚ mˈɑːlɪkjˌuːlz baɪ ðə dˈɑːnən ɪfˈɛkt.
At the same time the charges on the GAG chains attract ions and water molecules by the Donnan effect.