ɪn sɛptˈɛmbɚ nˈaɪntiːnhˈʌndɹɪd nˈaɪnti fˈoːɹ, dʒˈɛfɚsən fˈɜːst ɑːɹtˈɪkjʊlˌeɪɾᵻd ðə hˈɑːloʊdʒnˌoʊm θˈiəɹi ʌv ˌɛvəlˈuːʃən, æɾə pɹˌɛzəntˈeɪʃən æt kˈoʊld spɹˈɪŋ hˈɑːɹbɚ lˈæbɹətˌɔːɹi, æɾə sɪmpˈoʊsiəm "ɐ dˈɛkeɪd ʌv pˌiːsˌiːˈɑːɹ"
In September 1994, Jefferson first articulated the Hologenome Theory of Evolution, at a presentation at Cold Spring Harbor Laboratory, at a Symposium "A Decade of PCR"