ðə dʒˌiːˌeɪdʒˈiː tʃˈeɪnz ɑːɹ ˈeɪbəl tə fˈɔːɹm ˌæntɪpˈæɹəlˌɛl lˈɪŋks wɪð ˈʌðɚ dʒˌiːˌeɪdʒˈiː tʃˈeɪnz fɹʌm ɐdʒˈeɪsənt fˈɪbɹəlz, pɚhˈæps θɹuː ðə mˌiːdɪˈeɪʃən ʌv pˈɑːzᵻtˌɪvli tʃˈɑːɹdʒd ˈaɪɑːnz.
The GAG chains are able to form antiparallel links with other GAG chains from adjacent fibrils, perhaps through the mediation of positively charged ions.