ðeɪ ɑːɹ nˈoʊn fɹʌmðɪ ˌɔːɹɪnˈoʊkoʊ, ˈæmɐzˌɑːn, mˈæɡdeɪlnə, mˌæɹɐkˈeɪboʊ, pˌæɹənˈɑː, pɑːɹnˈeɪbə ænd sˈaʊ fɹænsˈɪskoʊ sˈɪstəmz.
They are known from the Orinoco, Amazon, Magdalena, Maracaibo, Paraná, Parnaíba and São Francisco systems.