æt ðə stˈɑːɹt ʌvðə sˈɛkənd wˈɜːld wˈɔːɹ hiː mˈuːvd tə ðə dᵻpˈɑːɹtmənt ʌv bˌaɪoʊkˈɛmɪstɹi æt ðə jˌuːnɪvˈɜːsᵻɾi ʌv kˈeɪmbɹɪdʒ wˌɛɹ hiː tˈʊk ˌɔn fɹˈɛd sˈændʒɚɹ æz hɪz pˌiːˈeɪtʃ dˈiː stˈuːdənt.
At the start of the Second World War he moved to the Department of Biochemistry at the University of Cambridge where he took on Fred Sanger as his PhD student.