fɔːɹ ˌoʊvɚ θˈɜːɾi jˈɪɹz hiː wʌzɐ baɪˈɑːlədʒi pɹəfˈɛsɚɹ æt wˈɑːʃɪŋtən jˌuːnɪvˈɜːsᵻɾi ænd ɐ dʒɪnˈɛɾɪks pɹəfˈɛsɚɹ æt wˈɑːʃɪŋtən jˌuːnɪvˈɜːsᵻɾi skˈuːl ʌv mˈɛdəsən.
For over 30 years he was a biology professor at Washington University and a genetics professor at Washington University School of Medicine.