hiː stˈʌdid nˈætʃɚɹəl sˈaɪənsᵻz æt ðə jˌuːnɪvˈɜːsᵻɾiz ʌv bɜːlˈɪn, wˈuːəzbˌɜːɡ ænd kˈiːl, ɹᵻsˈiːvɪŋ hɪz dᵻɡɹˈiː æt ðə lˈæɾɚɹ ˌɪnstɪtˈuːʃən ɪn wˈʌn θˈaʊzənd ˈeɪthˈʌndɹɪd ˈeɪɾi fˈoːɹ.
He studied natural sciences at the universities of Berlin, Würzburg and Kiel, receiving his degree at the latter institution in 1884.