dˈʊɹɹɪŋ ɐskəlˈɑːkjʊlɚ dɪvˈɛləpmənt pˈɑːkɪts lˈoʊkjuːlz fˈɔːɹm fˈɜːst wɪðˌɪn ðə vˈɛdʒɪtˌeɪɾɪv sˈɛlz ʌvðə fˈʌŋɡəs ænd ðˈɛn ˈɔːl ðə sˈʌbsᵻkwənt stɹˈʌktʃɚz fˈɔːɹm.
During ascolocular development pockets (locules) form first within the vegetative cells of the fungus and then all the subsequent structures form.