hiː lˈeɪɾɚ mˌeɪd ɪmpˈoːɹtənt dɪskˈʌvɚɹiz ɔnðɪ ˌɪntɚɹˈækʃən bᵻtwˌiːn ðə sˈaɪɾəskˌɛlᵻtən ænd ðə sˈɛl mˈɛmbɹeɪn, ɹɪzˈʌltɪŋ ˈɪn, ɐmˌʌŋ ˈʌðɚ θˈɪŋz, ðɪ aɪdˈɛntɪfɪkˈeɪʃən ʌvðə sˈaɪɾəskˌɛlᵻɾəl pɹˈoʊtiːnz vˈɪŋkjʊlˌɪn ænd tˈeɪlɪn.
He later made important discoveries on the interaction between the cytoskeleton and the cell membrane, resulting in, among other things, the identification of the cytoskeletal proteins vinculin and talin.