ˈæftɚ fˈɜːðɚ tɹˈeɪnɪŋ ɪnðə jˈuː.ˈɛs. pˈʌblɪk hˈɛlθ sˈɜːvɪs æt ðə nˈæʃənəl ˈɪnstɪtˌuːt ʌv ˈælɚdʒi ænd ɪnfˈɛkʃəs dɪzˈiːzᵻz, ænd hˈɑːɹvɚd mˈɛdɪkəl skˈuːl ɪn ɪnfˈɛkʃəs dɪzˈiːzᵻz, hiː dʒˈɔɪnd ðə jˌuːnɪvˈɜːsᵻɾi ʌv kˌælɪfˈoːɹniə, sˌæn dɪˈeɪɡoʊ ˈʌksd ɪn nˈaɪntiːnhˈʌndɹɪd sˈɛvənti sˈɪks, ˈʌltᵻmətli bᵻkˈʌmɪŋ dɪstˈɪŋɡwɪʃt pɹəfˈɛsɚɹ ʌv pæθˈɑːlədʒi ænd mˈɛdəsən, dᵻɹˈɛktɚɹ ʌvðə sˈɛntɚ fɔːɹ ˈeɪdz ɹᵻsˈɜːtʃ, dᵻɹˈɛktɚɹ ʌvðɪ ˌeɪtʃˌaɪvˈiː ˈɪnstɪtˌuːt ænd ðə flˈɔːɹəns sˈiːli ɹˈaɪfɚd tʃˈɛɹ ɪn ˈeɪdz ɹᵻsˈɜːtʃ.
After further training in the  U.S. Public Health Service at the National Institute of Allergy and Infectious Diseases, and Harvard Medical School in Infectious Diseases, he joined the University of California, San Diego (UCSD) in 1976, ultimately becoming Distinguished Professor of Pathology and Medicine, director of the Center for AIDS Research, director of the HIV Institute and the Florence Seeley Riford Chair in AIDS Research.