sˈɪstəmz stˈʌdid ɪŋklˈuːd dˌiːˌɛnˈeɪ ɹᵻpˈɛɹ, hˈoːɹmoʊnz, ɡɹˈoʊθ fˈæktɚz ænd hˈoːɹmoʊn slˈæʃ ɹᵻsˈɛptɚɹ ˌɪntɚɹˈækʃənz, sˈɛljʊlɚ sˈɪɡnəlɪŋ, kɹɪstˈælᵻnz lˈɛnz pɹˈoʊtiːnz, ɹᵻnˈɪn ænd ˌeɪtʃˌaɪvˈiː pɹˈoʊɾeɪs.
Systems studied include DNA repair, hormones, growth factors and hormone/receptor interactions, cellular signalling, crystallins (lens proteins), renin and HIV protease.