ðeɪ ɑːɹ ɹᵻlˈiːst wɛn mətʃˈʊɹ baɪ ðə dˌɪsˌɪntᵻɡɹˈeɪʃən ʌvðə spɔɹˈæŋɡiəm wˈɔːl, ɔːɹ æz ɐ hˈoʊl spˈɔɹɐŋɡɪˌoʊl ðæt sˈɛpɚɹˌeɪts fɹʌmðə spˈɔɹɐŋɡɪˌɑːfoːɹ.
They are released when mature by the disintegration of the sporangium wall, or as a whole sporangiole that separates from the sporangiophore.