æz ɐ pˈoʊstdəktɚɹəl ɹᵻsˈɜːtʃɚ hiː wˈɜːkt æt ðə plˈænt bɹˈiːdɪŋ ˈɪnstɪtˌuːt ɪn kˈeɪmbɹɪdʒ, ˈɪŋɡlənd: ðɛɹ hiː ɐdˈæptᵻd ðə dʒˌiːjˌuːˈɛs ɐsˈeɪ fɚðə jˈuːs ɪn plˈænts.
As a postdoctoral researcher he worked at the Plant Breeding Institute in Cambridge, England: there he adapted the GUS assay for the use in plants.