Nikolaas Tinbergen shared the 1973 Nobel in Medicine with the zoologist who was of what nationality?
Nikolaas Tinbergen Nikolaas 'Niko' Tinbergen {'1': ", '2': ", '3': ", '4': "} ( ; ] ; 15 April 1907 – 21 December 1988) was a Dutch biologist and ornithologist who shared the 1973 Nobel Prize in Physiology or Medicine with Karl von Frisch and Konrad Lorenz for their discoveries concerning organisation and elicitation of individual and social behaviour patterns in animals. He is regarded as one of the founders of modern ethology, the study of animal behaviour.
Jakob von Uexküll Jakob Johann Baron von Uexküll (] ; 8 September 1864 – 25 July 1944) was a Baltic German biologist who worked in the fields of muscular physiology, animal behaviour studies, and the cybernetics of life. However, his most notable contribution is the notion of "Umwelt", used by semiotician Thomas Sebeok and philosopher Martin Heidegger. His works established biosemiotics as a field of research.