Fischer (2004) considered the family to consist of three subfamilies – Antirrhinoideae, Gratioloideae, and Digitalidoideae. He further divided the Gratioloideae into five tribes – Gratioleae, Angeloniaeae, Stemodieae, Limoselleae, and Lindernieae. He then divided the Gratioleae, with its 16 genera (and about 182 species) into three subtribes – Caprarinae, Dopatrinae, and Gratiolinae. The Gratiolinae had 10 genera (about 121 species) distributed through temperate and tropical America – Bacopa and Mecardonia (formerly Herpestis), Amphianthus, Gratiola, Sophronanthe, Benjaminia, Scoparia, Boelkea, Maeviella, and Braunblequetia. Many of these were transferred to the family Plantaginaceae, in the tribe Gratioleae.
fischer (2004) considered the family to consist of three subfamilies – antirrhinoideae gratioloideae and digitalidoideae he further divided the gratioloideae into five tribes – gratioleae angeloniaeae stemodieae limoselleae and lindernieae he then divided the gratioleae with its 16 genera (and about 182 species) into three subtribes – caprarinae dopatrinae and gratiolinae the gratiolinae had 10 genera (about 121 species) distributed through temperate and tropical america – bacopa and mecardonia (formerly herpestis) amphianthus gratiola sophronanthe benjaminia scoparia boelkea maeviella and braunblequetia many of these were transferred to the family plantaginaceae in the tribe gratioleae