In addition to the harmony found in suffixes, there is a harmony restriction on word stems where in stems with more than one syllable all vowels are required to be of the same lip rounding and tongue height dimensions. For example, a stem must contain all high rounded vowels or all low rounded vowels, etc. This restriction is further complicated by (i) long high vowels being lowered and (ii) an epenthetic vowel which does not harmonize with stem vowels.
In addition to the harmony found in suffixes, there is a harmony restriction on word stems where in stems with more than one syllable all vowels are required to be of the same lip rounding and tongue height dimensions. For example, a stem must contain all high rounded vowels or all low rounded vowels, etc. This restriction is further complicated by (i) long high vowels being lowered and (ii) an epenthetic vowel which does not harmonize with stem vowels.