title: ImportError: cannot import name signals when importing wsgi file in Django 1.10
body: <p>I'm getting the following error trying to run Django using apache:</p>
  
  <pre><code>mod_wsgi (pid=3294): Target WSGI script '/www/cocurate2_dev/startup.wsgi' cannot be loaded as Python module.
  mod_wsgi (pid=3294): Exception occurred processing WSGI script '/www/cocurate2_dev/startup.wsgi'.
  Traceback (most recent call last):
    File "/www/cocurate2_dev/startup.wsgi", line 3, in &lt;module&gt;
      from django.core.wsgi import get_wsgi_application
    File "/www/cocurate2_dev/components/lib/python2.7/site-packages/django/core/wsgi.py", line 2, in &lt;module&gt;
      from django.core.handlers.wsgi import WSGIHandler
    File "/www/cocurate2_dev/components/lib/python2.7/site-packages/django/core/handlers/wsgi.py", line 10, in &lt;module&gt;
      from django import http
    File "/www/cocurate2_dev/components/lib/python2.7/site-packages/django/http/__init__.py", line 5, in &lt;module&gt;
      from django.http.response import (
    File "/www/cocurate2_dev/components/lib/python2.7/site-packages/django/http/response.py", line 13, in &lt;module&gt;
      from django.core.serializers.json import DjangoJSONEncoder
    File "/www/cocurate2_dev/components/lib/python2.7/site-packages/django/core/serializers/__init__.py", line 23, in &lt;module&gt;
      from django.core.serializers.base import SerializerDoesNotExist
    File "/www/cocurate2_dev/components/lib/python2.7/site-packages/django/core/serializers/base.py", line 4, in &lt;module&gt;
      from django.db import models
    File "/www/cocurate2_dev/components/lib/python2.7/site-packages/django/db/models/__init__.py", line 4, in &lt;module&gt;
      from django.db.models import signals  # NOQA
  ImportError: cannot import name signals
  </code></pre>
  
  <p>Everything works find when using manage.py runserver. I've updated the wsgi file to be the same as the default one from django 1.10.
  I've checked that signals.py exists inside django, I've also cleaned .pyc files.</p>