Did Pain control the animals from his Summoning jutsu?


I'm a fan of _Naruto_ , I wonder if Pain was manipulating his summoned beasts,
as I noticed rods in all the animals. Also, were those animals dead?




Yes! According to wiki page [here](https://naruto.fandom.com/wiki/Animal_Path)
This is a list Pain can summon :

> The Animal Path (畜生道, Chikushōdō) grants a Rinnegan user the ability of
> summoning various animals and creatures to aid it in battle. These creatures
> appear to be immortal, as they were killed and later summoned again
>
>   * Giant Drill-Beaked Bird
>   * Giant Multi-Headed Dog
>   * Giant Snake-Tailed Chameleon
>   * Giant Panda
>   * Giant Ox
>   * Giant Rhino
>   * Giant Crustacean
>   * Giant Centipede
>




Yes! According to wiki page [here](https://naruto.fandom.com/wiki/Animal_Path)
This is a list Pain can summon :

> The Animal Path (畜生道, Chikushōdō) grants a Rinnegan user the ability of
> summoning various animals and creatures to aid it in battle. These creatures
> appear to be immortal, as they were killed and later summoned again
>
>   * Giant Drill-Beaked Bird
>   * Giant Multi-Headed Dog
>   * Giant Snake-Tailed Chameleon
>   * Giant Panda
>   * Giant Ox
>   * Giant Rhino
>   * Giant Crustacean
>   * Giant Centipede
>

