grin grin-full-closed grin-full-open grin-low-closed grin-low-open groom groom-branch-clasp groom briefly groom by hand groom carcass groom, dyadic groom-hand-clasp groom-hand-clasp unilaterally groom leaf groom mutually groom object or substrate groom, polyadic groom reciprocally groom self groom unilaterally Appendix Toshisada Nishida Toshisada Nishida Toshisada Nishida Toshisada Nishida Toshisada Nishida Takahisa Matsusaka Toshisada Nishida Koichiro Zamma Toshisada Nishida Toshisada Nishida Koichiro Zamma Toshisada Nishida Koichiro Zamma Toshisada Nishida Toshisada Nishida Toshisada Nishida Toshisada Nishida Toshisada Nishida Toshisada Nishida Toshisada Nishida Toshisada Nishida Toshisada Nishida Toshisada Nishida Toshisada Nishida Toshisada Nishida Toshisada Nishida Toshisada Nishida Koichiro Zamma Toshisada Nishida Toshisada Nishida Toshisada Nishida Toshisada Nishida Toshisada Nishida Toshisada Nishida Toshisada Nishida Toshisada Nishida Koichiro Zamma Koichiro Zamma Toshisada Nishida Toshisada Nishida Toshisada Nishida Koichiro Zamma Toshisada Nishida Toshisada Nishida Appendix groom with mouth