supine ride ventral ride ventral with limb extended 241 Toshisada Nishida Koichiro Zamma Toshisada Nishida Toshisada Nishida Toshisada Nishida Toshisada Nishida Koichiro Zamma Toshisada Nishida Takahisa Matsusaka Toshisada Nishida Toshisada Nishida Koichiro Zamma Toshisada Nishida Koichiro Zamma Toshisada Nishida Koichiro Zamma Koichiro Zamma Toshisada Nishida Toshisada Nishida Toshisada Nishida Toshisada Nishida Toshisada Nishida Koichiro Zamma Toshisada Nishida Toshisada Nishida Koichiro Zamma Toshisada Nishida Toshisada Nishida Koichiro Zamma Toshisada Nishida Toshisada Nishida Toshisada Nishida Koichiro Zamma Koichiro Zamma Takahisa Matsusaka Toshisada Nishida Koichiro Zamma Koichiro Zamma Koichiro Zamma Toshisada Nishida Koichiro Zamma Koichiro Zamma Koichiro Zamma Toshisada Nishida 242 rock back and forth rock side to side roll rotate fruit rub dorsum rub dorsum to conspecific rub genitals rub genitals to substrate rub hand or foot rub hand with