beg clamp elbow extend hand to beg fend give mouth for begging peer push away share food take turn around twist 2-4. Feeding and foraging: Hunting chase drag to kill dunk eat meat expel hunt hunt with tool knock with both arms knock with one arm monitor monkeys scrub pelt share food stalk twist wash wash colobus skin 3-1. Expressive behavior: Vocalization bark bark, infantile choke in tantrum clack teeth cough bark distress call grunt grunt, aha grunt, extended grunt, food grunt in bed hoo hoot huu mass excitement pant pant-bark pant-grunt pant-hoot pant in copulation play-pant scream scream, food smack lip sputter squeal in copulation staccato call vocalize waa bark whimper whimper-scream wraa 3-2. Expressive behavior: Facial expression compress lips flip lip funny face grin grin-full-closed grin-full-open