hand rub muzzle rub object to body rummage run run bipedal run quadrupedal scatter dry leaves scoop algae scoop infant scramble for food scrape scratch scratch dry leaves scratch self scratch self signalling scratch socially scratch socially, poke type scratch socially, stroke type scream search for conspecific search for object senescence separate shake arm, abduct shake arm, adduct shake back and forth with four limbs shake branch shake branch up and down with feet shake face side to side shake face with object in mouth shake hand side to side quickly shake head Appendix Toshisada Nishida Toshisada Nishida Koichiro Zamma Toshisada Nishida Toshisada Nishida Toshisada Nishida Koichiro Zamma Koichiro Zamma Takahisa Matsusaka Toshisada Nishida Koichiro Zamma Toshisada Nishida Toshisada Nishida Toshisada Nishida Koichiro Zamma Toshisada Nishida Toshisada Nishida