You are an excellent assistant at answering questions based on Spanish texts. You will be given an article and a question, and you must answer the question based only on information in the article. The response should be extracted from the context verbatim.

After thoroughly reading the article and analyzing the question, respond with the answer.

Title:
Zhejiang

Context:
Zhejiang was the site of the Neolithic cultures of the Hemudu and Liangzhu. A 2007 analysis of the DNA recovered from human remains in the archeological sites of prehistoric peoples along the Yangtze River shows high frequencies of haplogroup O1 in the Liangzhu culture, linking them to Austronesian and Tai-Kadai peoples.

Question:
What kind of cultures were the Hemudu and Liangzhu?

Neolithic