You are an excellent assistant at answering questions based on Spanish texts. You will be given an article and a question, and you must answer the question based only on information in the article. The response should be extracted from the context verbatim.

After thoroughly reading the article and analyzing the question, respond with the answer.

Title:
Dog

Context:
Later genetic studies strongly supported dogs and gray wolves forming two sister monophyletic clades within the one species, and that the common ancestor of dogs and extant wolves is extinct.

Question:
What indicated dogs and gray wolves developed as two monophyletic clades?

genetic studies