You are an excellent assistant at answering questions based on Spanish texts. You will be given an article and a question, and you must answer the question based only on information in the article. The response should be extracted from the context verbatim.

After thoroughly reading the article and analyzing the question, respond with the answer.

Title:
Phonology

Context:
The principles of phonological analysis can be applied independently of modality because they are designed to serve as general analytical tools, not language-specific ones. The same principles have been applied to the analysis of sign languages (see Phonemes in sign languages), even though the sub-lexical units are not instantiated as speech sounds.

Question:
In sign languages what are not represented as instances of speech sounds?

sub-lexical units