[PAR] [TLE] Tim Hunt [SEP] Sir Richard Timothy Hunt, (born 19 February 1943) is a British biochemist and molecular physiologist.  He was awarded the 2001 Nobel Prize in Physiology or Medicine with Paul Nurse and Leland H. Hartwell for their discoveries of protein molecules that control the division of cells.  In particular, Hunt discovered what he called cyclin: a protein in fertilised sea urchin eggs which cyclically aggregates and is depleted during cell division cycles. [PAR] [TLE] David Colquhoun [SEP] David Colquhoun, {'1': ", '2': ", '3': ", '4': "} (born 19 July 1936) is a British pharmacologist at University College London (UCL).  He has contributed to the general theory of receptor and synaptic mechanisms, and in particular the theory and practice of single ion channel function.  He previously held the A.J. Clark chair of Pharmacology at UCL, and was the Hon. Director of the Wellcome Laboratory for Molecular Pharmacology.  He was elected a Fellow of the Royal Society (FRS) in 1985 and an honorary fellow of UCL in 2004.  Colquhoun runs the website "DC's Improbable Science", which is critical of pseudoscience, particularly alternative medicine, and managerialism.  He was critical of Tim Hunt following the controversies of the 2015 World Conference of Science Journalists in Seoul.
[PAR] [TLE] Tim Hunt [ SEP ] Sir Richard Timothy Hunt , ( born 19 February 1943 ) is a British biochemist and molecular physiologist . He was awarded the 2001 Nobel Prize in Physiology or Medicine with Paul Nurse and Leland H. Hartwell for their discoveries of protein molecules that control the division of cells . In particular , Hunt discovered what he called cyclin : a protein in fertilised sea urchin eggs which cyclically aggregates and is depleted during cell division cycles . [PAR] [TLE] David Colquhoun [ SEP ] David Colquhoun , { ' 1 ' : " , ' 2 ' : " , ' 3 ' : " , ' 4 ' : " } ( born 19 July 1936 ) is a British pharmacologist at University College London ( UCL ) . He has contributed to the general theory of receptor and synaptic mechanisms , and in particular the theory and practice of single ion channel function . He previously held the A.J. Clark chair of Pharmacology at UCL , and was the Hon . Director of the Wellcome Laboratory for Molecular Pharmacology . He was elected a Fellow of the Royal Society ( FRS ) in 1985 and an honorary fellow of UCL in 2004 . Colquhoun runs the website " DC 's Improbable Science " , which is critical of pseudoscience , particularly alternative medicine , and managerialism . He was critical of Tim Hunt following the controversies of the 2015 World Conference of Science Journalists in Seoul .
After the controversies of the 2015 World Conference of Science Journalists in Seoul, David Colquhoun was critical of a British biochemist who was awarded the Nobel Prize in Physiology or Medicine in what year?
After the controversies of the 2015 World Conference of Science Journalists in Seoul , David Colquhoun was critical of a British biochemist who was awarded the Nobel Prize in Physiology or Medicine in what year ?
2001
2001