[PAR] [TLE] Nicaragua [SEP] Nicaragua ( ; ] ), officially the Republic of Nicaragua (Spanish:    ), is the largest country in the Central American isthmus, bordered by Honduras to the north, the Caribbean to the east, Costa Rica to the south, and the Pacific Ocean to the west.  Nicaragua's capital, Managua, is the country's largest city and the third-largest city in Central America.  The multi-ethnic population of six million includes indigenous peoples, Europeans, Africans, and Asians.  The main language is Spanish.  Native tribes on the eastern coast speak their own languages. [PAR] [TLE] Aratus pisonii [SEP] Aratus pisonii, commonly known as the mangrove tree crab, is a species of crab which lives in mangrove trees in tropical and subtropical parts of the Americas, from Florida to Brazil on the Atlantic coast, and from Nicaragua to Peru on the Pacific coast.  It feeds mostly on the leaves of the mangroves, but is an omnivore, and prefers animal matter when possible.  "A. pisonii" is the only species in the monotypic genus Aratus.  The specific epithet "pisonii" commemorates the Dutch naturalist Willem Piso who travelled in Brazil in 1638 with Georg Marggraf.
[PAR] [TLE] Nicaragua [ SEP ] Nicaragua ( ; ] ) , officially the Republic of Nicaragua ( Spanish : ) , is the largest country in the Central American isthmus , bordered by Honduras to the north , the Caribbean to the east , Costa Rica to the south , and the Pacific Ocean to the west . Nicaragua 's capital , Managua , is the country 's largest city and the third - largest city in Central America . The multi - ethnic population of six million includes indigenous peoples , Europeans , Africans , and Asians . The main language is Spanish . Native tribes on the eastern coast speak their own languages . [PAR] [TLE] Aratus pisonii [ SEP ] Aratus pisonii , commonly known as the mangrove tree crab , is a species of crab which lives in mangrove trees in tropical and subtropical parts of the Americas , from Florida to Brazil on the Atlantic coast , and from Nicaragua to Peru on the Pacific coast . It feeds mostly on the leaves of the mangroves , but is an omnivore , and prefers animal matter when possible . " A. pisonii " is the only species in the monotypic genus Aratus . The specific epithet " pisonii " commemorates the Dutch naturalist Willem Piso who travelled in Brazil in 1638 with Georg Marggraf .
What is the capital city of this Central American isthmus, to which the species of crab Aratus pisonii is native, bordering with Costa Rica?
What is the capital city of this Central American isthmus , to which the species of crab Aratus pisonii is native , bordering with Costa Rica ?
Managua
Managua