original_query: What were the characteristics of the petiole and stem morphology that were analyzed in the study?
query: What were the characteristics of the petiole and stem morphology of Lophosoria quadripinnata that were analyzed in the study?
answer: The study analyzed characteristics of the petiole and stem morphology that hypothesized a close relationship with Metaxyaceae.  However, subsequent analyses suggested these families were not related.