anchor.diff:  bongardii var. pisciodora (Donadini & Riousset) Kuyper,Leiospora,Regular | Subphaseoliform,Obtuse,[12-16.7],[6.5-8.7],[1.7-2.1],Smooth,Claviform,[37-59],[10-14],[4-4],Yes,Cylindrical | Claviform | Utriform,[33-77],[8-22],No,[0.5-1.5],n/a,n/a,n/a,n/a,n/a,No,n/a,n/a,n/a,No,n/a,n/a,Yes,Cylindrical | Claviform | Utriform,n/a,n/a,No,n/a,n/a,No,n/a,n/a,n/a,No,n/a,n/a,n/a,n/a,Soil,Oak | Spruce | Pine,[15-42],[15-42],Attenuate,n/a,n/a,Pelargonium | Fishy | Not distinctive,Inosperma,Cervicolores,,,"1) Vernal variants from the Mediterranean Region are generally more robust than summer and autumnal variants from West and Central Europe. No taxonomic value can be given to this difference, however. 2) Mr J. C. Donadini affirmed in a letter that I. pisciodora cannot be considered an autonomous species, but is only an infraspecific taxon of I. bongardii. 3) Although nothing is known about the composition of the olfactory substances of var. bongardii and var. pisciodora, I consider it very likely that they are chemically much related. Young specimens of var. pisciodora often possess a mixture of a Pelargonium-smell and a smell of overripe pears. Both smells can also be observed in I. calamistrata (Fr. : Fr.) Gillet.","Kuyper, Thomas W. 1986. A revision of the genus Inocybe in Europe : I. subgenus inosperma and the smooth-spored species of subgenus Inocybe, Rijksherbarium, Leiden, 247 p.; p. 41-42",,,,
   borealis M. Lange,Goniospora,Nodulose | Angular,Subconical | Conical,[7.2-10],[5-6.9],[1.3-1.9],Sometimes with a strongly protruding apical nodule,Claviform,[24-40],[9-13],[2-4],Yes,Lageniform | Cylindrical | Fusiform | Metuloid,n/a,n/a,Yes | No,n/a,n/a,n/a,n/a,n/a,n/a,Yes,Lageniform | Cylindrical | Fusiform | Metuloid,[41-89],[12-26],Yes | No,[2.5-3],n/a,Yes,Lageniform | Cylindrical | Fusiform | Metuloid,n/a,n/a,Yes | No,[1-2],n/a,No,n/a,n/a,n/a,No,n/a,n/a,Yes,n/a,Sandy soil,Spruce | Fir | Birch,[13-45],[13-45],Equal | Widening slightly downwards,n/a,n/a,Acidulous | Spermatic | Not distinctive,Clypeus,,,,"Inocybe borealis is a variable species in many aspects. Our identifications of boreal collections, a subarctic collection, and I. heterochrominea are primarily based on ITS and LSU sequences, because the spores diverged so much from the I. borealis type. They were consistently smaller (except in KK141/03), paler, and more strongly nodulose. The Nordic ITS sequences were identical with the type sequence of I. borealis in two specimens (KK190/08 - GenBank JN580813, KK&JV27051F - JN580814), or deviated from the type by one gap (JV9244 - JN580815), by one common base in nine specimens (KK&JV26120F - JN580816), by one other base in JV15250F (JN580817), by one base and a gap of 12 bases (KK141/03 - JN580818), or by two bases (Heinonen 307-95 - JN580819). The I. heterochrominea type deviated from the I. borealis type by one base and a gap of one base, when 42 bases were lacking in the middle of the I. heterochrominea sequence and three loci with ambiguous bases were ignored. The LSU sequences of the type of I. borealis and a collection with one different base in ITS (KK&JV26120F - JN580889) diverged by one base as well. An arctic collection from Svalbard (Online Resource Fig. S3) resembled the Greenland type in its spores, the presence of velipellis (absent in the other collections), and identical ITS (in the phylogram, JN580820). These specimens thus seem to form their own group. They both also have dark brown pilei seemingly unlike most boreal collections (the colours may have changed in exsiccata). Still, the two boreal collections with the type ITS also have dark brown pilei (Online Resource Fig. S2), but in spores they resemble other boreal specimens rather than the type; thus forming an intermediate group. Overall, the strength of nodularity varies rather much among the boreal specimens, and rarely the spores are elongated (KK141/03) as in the type of Inocybe borealis. We were unable to find more distinct morphological boundaries between the specimens differing in ITS or between arctic and boreal/subarctic collections, and thus we tentatively determine them all to be the same very variable species. The observed morphological differences may have been caused by environment, or may be genetically intraspecific. The amount of velipellis, for example, often varies within species and seems to increase in arctic and alpine environments. The microscopic and macroscopic features of Inocybe heterochrominea agreed with our Nordic collections of I. borealis. Accordingly, we have reduced it to a synonym. The walls of the cystidia of the I. heterochrominea holotype (a piece of a pileus) seemed thinner than in the drawings of Grund and Stuntz (1983), and the spores did not seem quite as strongly nodulose. I. heterochrominea was described from mixed and conifer forests of Michigan, Nova Scotia, and Washington. In Finland, I. borealis grows in various habitats, and we assume Betula to be one host. Lange (1957) did not mention any plant species from the type site, but he described the vegetation and fungi of the northern slopes, for example, stating that Betula nana was the dominant shrub and Salix glauca was scattered. Inocybe acutoides is a closely related sibling species associated with Salix. See our comments regarding I. acutoides, and the diagnostic key.","Kokkonen, Katri; Vauras, Jukka. Eleven new boreal species of Inocybe with nodulose spores, Mycol Progress (2012) 11:299-341; p. 309-312",,,,
   "boreocarelica Kokkonen & Vauras, sp. nov.",Goniospora,Strongly nodulose,Subconical | Conical,[7-10.1],[5-7.5],[1.2-1.7],Nodules usually small,Claviform,[24-38],[8-12],[4-4],Yes,Lageniform | Ellipsoid | Metuloid,n/a,n/a,Yes | No,n/a,n/a,Yes,n/a,n/a,n/a,Yes,Lageniform | Metuloid,[45-82],[11-26],Yes,[2-4],n/a,Yes,Lageniform | Fusuform | Cylindrical | Claviform | Metuloid,n/a,n/a,Yes | No,[1-2],n/a,No,n/a,n/a,n/a,No,n/a,n/a,Yes,n/a,Soil,Spruce | Birch | Alder | Heath forest mainly,[14-35],[14-35],Tapering | Roundish,n/a,n/a,Acidulous | Not distinctive,Clypeus,,,,"Inocybe boreocarelica may be difficult to identify, but it is characterised by its reddish brown, flocculose stipe and rather small spores. It may resemble I. borealis both in morphology and habitat. Both sometimes have a yellow brown pileus cracked into scales at umbo, as observed in an old cemetery (Pitkamaki, Nurmes) where they grow near each other. However, the cystidial walls of I. boreocarelica are often distinctly thickened in the neck. Inocybe boreocarelica also resembles I. bufonia microscopically, but the spores of I. bufonia are more often elongated. I. bufonia differs more distinctly in its darker appearance and moister habitat. The ITS sequence of the Norwegian specimen deviated from the others by one base (GenBank JN580835).","Kokkonen, Katri; Vauras, Jukka. Eleven new boreal species of Inocybe with nodulose spores, Mycol Progress (2012) 11:299-341; p. 314-315",,,,
  -bresadolae Massee,Goniospora,,,,,,,,,,,,,,,,,,,,,,,,,,,,,,,,,,,,,,,,,,,,,,,,,,,,,Clypeus,Marginatae,Rubellae,,,,,,,
  +bresadolae Massee,Goniospora,Tuberculate | Polygonale,Subconical | Conical,[7-9],[5-6],n/a,"With about 6 to 8 rather prominent, rounded nodules",Claviform,n/a,n/a,[4-4],Yes,n/a,n/a,n/a,n/a,n/a,n/a,n/a,n/a,n/a,n/a,Yes,Variables | Metuloid,[40-75],[12-23],Yes | Sparse | No,n/a,n/a,No,n/a,n/a,n/a,No,n/a,n/a,No,n/a,n/a,n/a,No,n/a,n/a,n/a,n/a,Soil,Conifers | Hardwoods,[20-40],[20-40],Marginate bulbous | Submarginate bulbous,n/a,n/a,Pears,Clypeus,Marginatae,Rubellae,n/a,"Species innocua videtur, vermutamen ne colligenda quidem ut edatur, quia cum aliis similibus facile confusa.","Bresadola, Ab. Dr. J. ; Alessio, C. L. Iconographia Mycologica, Vol. XXIX, Inocybe. Societa Botanica Italiana, 1980",,,,
   brevicystis Metrod ex Kuyper,Leiospora,Subamygdaliform,Obtuse | Subconical,[9-11],[5-6.5],[1.5-1.8],Smooth,Claviform,[27-36],[9-11],[4-4],Yes,Cylindrical | Claviform | Subfusiform | Metuloid,n/a,n/a,Yes,n/a,n/a,Yes,Claviform,n/a,n/a,Yes,Cylindrical | Claviform | Subfusiform | Metuloid,[31-51],[9-14],Yes,[1-2],n/a,Yes,Cylindrical | Claviform | Subfusiform | Metuloid,n/a,n/a,n/a,n/a,n/a,Yes,n/a,n/a,n/a,n/a,n/a,n/a,n/a,n/a,Soil,Oak,[20-30],[20-30],Not bulbous,n/a,n/a,Subspermatic,Inocybe,,,,"The macroscopical description has been based on Metrod (in Bull. trimest. Soc. mycol. Fr. 72 : 126. 1956) and Reumaux (in Does mycol. 14 (54-55) : 5. 1984). 2) The rather short and narrow cystidia are indicative for I. brevicystis. Inocybe mycenoides Kuyp. posses the same cystidia, but that species differs in having a mycenoid habit and more yellow lamellae. Inocybe ochroalba Bruylants might show some macroscopical resemblance to I. brevicystis, but differs in cystidial characters and having a distinctly conical spore apex.","Kuyper, Thomas W. 1986. A revision of the genus Inocybe in Europe : I. subgenus inosperma and the smooth-spored species of subgenus Inocybe, Rijksherbarium, Leiden, 247 p.; p. 212-213",,,,
   brunneolipes Grund & D.E. Stuntz,Leiospora,Inequilaterally | Amygdaliform,Obtuse | Subconical,[7-10],[4.5-6],n/a,Smooth,Claviform,[22-30],[8-10],[2-4],Yes,Fusoid | Claviform | Metuloid,[16-36],[9-15],Yes | No,n/a,n/a,n/a,n/a,n/a,n/a,Yes,Fusoid | Sublanceolate | Metuloid,[45-68],[11-19],Yes,[3-4],n/a,Yes,Fusoid | Sublanceolate | Metuloid,[45-90],[9-16],Yes | No,n/a,n/a,Yes,Fusoid | Sublanceolate | Metuloid,[40-90],[10-22],Yes | No,n/a,n/a,n/a,n/a,Sandy soil,Conifers | Spruce | Fir,[20-35],[20-35],Equal,n/a,jacobi,Spermatic,,,,," The distinctive features of this Inocybe are the smooth umbrinous pileus, paler brown pruinose stipe, smooth moderate-sized spores, and thick-walled, hyaline pleurocystidia. I. brunneolipes takes its place in a small group of similar taxa, that includes I. reducta J. Lange and I. descissa f. bisporigera J. Lange. The former differs from I. brunneolipes by smaller spores and bulbous-based stipe, the latter species is distinct by 2-spored basidia, with all spores 10-12(-13) x 5-6 micron, and in the smaller size and much more slender stipe. In the collection of I. brunneolipes from Washington 2-spored basidia predominate and most of the spores are 11-14 x 6-7 micron, whereas in the Nova Scotian collection 2-spored basidia are decidedly in the minority and the smaller spores predominate. In all other respects the specimens of the two collections are practically indistinguishable.","Grund, D.W.; Stuntz, D.E. Nova Scotian Inocybes II., Mycologia, vol. 62, 1970; p. 928-929",,,,
   brunneotomentosa Huijsman,Leiospora,Subamygdaliform,Subconical,[7.5-9.5],[4.5-5.5],[1.5-1.8],Smooth,Claviform,[23-29],[7-9],[4-4],Yes,Claviform | Cylindrical | Metuloid,n/a,n/a,Yes,n/a,n/a,Yes,Claviform,n/a,n/a,Yes,Claviform | Cylindrical | Metuloid,[34-52],[14-23],Yes,[2-2.5],n/a,Yes,Claviform | Cylindrical | Metuloid,n/a,n/a,n/a,n/a,n/a,No,n/a,n/a,n/a,No,n/a,n/a,n/a,n/a,Soil,Oak | Beech | Blue-beech,[15-40],[15-40],Equal,n/a,n/a,Spermatic,Inocybe,,,,"The macroscopical description has been copied from Huijsman (in Persoonia) 9 : 473. 1978). Inocybe brunneotomentosa can be confused with two other species with a darkening stipe, viz. I. melanopus Stuntz and I. tenebrosa Quel. The former species has a rather pale ochraceous pileus, and differently-shapped, rather thin-walled pleurocystidia, whereas the latter species lacks a cortina and has the stipe covered with caulocystidia throughout.","Kuyper, Thomas W. 1986. A revision of the genus Inocybe in Europe : I. subgenus inosperma and the smooth-spored species of subgenus Inocybe, Rijksherbarium, Leiden, 247 p.; p. 127-128",,,,
  
positive.code: sororia Kauffman,Leiospora,Ellipsoid | Elongate-ellipsoid | Subinequilateral,Obtuse,[9-16],[5.5-8],n/a,Smooth,Claviform,n/a,n/a,n/a,n/a,n/a,n/a,n/a,n/a,n/a,n/a,n/a,n/a,n/a,n/a,No,n/a,n/a,n/a,No,n/a,n/a,n/a,n/a,n/a,n/a,n/a,n/a,n/a,n/a,n/a,n/a,n/a,n/a,n/a,n/a,n/a,n/a,Ground,Hardwoods | Conifers,[20-70],[20-70],Subbulbous,n/a,n/a,Green-corn | Not distinctive,,,,,n/a,"Kauffman, Calvin Henry. North American Flora, vol. 10(4), The New York Botanical Garden, 1924; p. 259, no 100",,,,
  splendens var. phaeoleuca (Kuehner) Kuyper,Leiospora,Subamygdaliform | Amygdaliform | Suprahilar depression,Subconical,[8-11.5],[5-6.5],[1.4-1.9],Smooth,Claviform,[24-30],[8-11],[2-4],Yes,Fusiform | Utriform | Claviform | Sublageniform | Metuloid,n/a,n/a,Yes,n/a,n/a,Yes,Spheropedunculate | Pyriform,n/a,n/a,Yes,Fusiform | Utriform | Claviform | Sublageniform | Metuloid,[49-78],[12-22],Yes,[2-2.5],n/a,Yes,Fusiform | Utriform | Claviform | Sublageniform | Metuloid,n/a,n/a,n/a,n/a,n/a,Yes,Fusiform | Utriform | Claviform | Sublageniform | Metuloid,n/a,n/a,n/a,n/a,n/a,Yes,n/a,Calcereous sand | Loam,Beech | Oak | Blue-beech | Birch | Poplar | Alder | Willow | Pine,[12-46],[12-46],Equal | Subbulbous,n/a,n/a,Faint | Pelargonium | Subspermatic | Spermatic,Inocybe,,,,"Although the extreme variants of I. splendens, I. terrifera, and I. phaeoleuca are very different, they are united here as there are no microscopical differences between thse taxa, and the macroscopical differences are at least partly phenotypically determined. Besides, several locally adapted ecotypes have been found. Recognition on specific level would ultimately result in a rather large number of species, the majority of these with a local distribution, but such a taxonomy would hardly possess any predictive value. Considering the above, it appears unavoidable to accep only one species with two rather well-delimited varieties.","Kuyper, Thomas W. 1986. A revision of the genus Inocybe in Europe : I. subgenus inosperma and the smooth-spored species of subgenus Inocybe, Rijksherbarium, Leiden, 247 p.; p. 215-216",,,,
  splendens var. splendens R. Heim,Leiospora,Subamygdaliform | Amygdaliform | Suprahilar depression,Subconical,[8-12],[5-7],[1.5-1.9],Smooth,Claviform,[25-38],[8-12],[4-4],Yes,Claviform | Fusiform | Utriform | Metuloid,n/a,n/a,Yes,n/a,n/a,Yes,Pyriform,n/a,n/a,Yes,Claviform | Fusiform | Utriform | Metuloid,[49-82],[14-29],Yes,[2-3],n/a,Yes,Claviform | Fusiform | Utriform | Metuloid,n/a,n/a,n/a,n/a,n/a,Yes,Claviform | Fusiform | Utriform | Metuloid,n/a,n/a,n/a,n/a,n/a,Yes,n/a,Calcereous sand | Clay,Beech | Oak | Basswood | Birch | Poplar | Alder | Willow,[23-78],[23-78],Equal | Submarginately | Marginately bulbous,n/a,n/a,Pelargonium | Spermatic,Inocybe,,,,"Although the extreme variants of I. splendens, I. terrifera, and I. phaeoleuca are very different, they are united here as there are no microscopical differences between thse taxa, and the macroscopical differences are at least partly phenotypically determined. Besides, several locally adapted ecotypes have been found. Recognition on specific level would ultimately result in a rather large number of species, the majority of these with a local distribution, but such a taxonomy would hardly possess any predictive value. Considering the above, it appears unavoidable to accep only one species with two rather well-delimited varieties.","Kuyper, Thomas W. 1986. A revision of the genus Inocybe in Europe : I. subgenus inosperma and the smooth-spored species of subgenus Inocybe, Rijksherbarium, Leiden, 247 p.; p. 215-216","1) The collection from France is characterised by the complete lack of pleurocystidia and the extreme scarcety of cheilocystidia. More material of this variant seems needed for a better evaluation of the relevance of this character state difference. 2) Inocybe splendens var. splendens is rather variable is outward aspect and in colour of the pileus. Specimens with a well-development subvicid velipellis are rather pale and bear conspicuous clods of earth, whereas specimens with a weakly developed, more innate velipellis are much darker (even to dark brown) and have a smooth pileus. The former type is expecially encountered on clayey soils and seems somewhat more xerophytic. This type probably consists of several locally adapted ecotypes. 3) Considering the rather robust specimens as depicted by Furrer-Ziogas (in Schweiz. Z. Pilzk. 43: pl. 1, f. 5-7, 1965) and Stangl (in Z. Pilzk. 37: pl. 7. 1971). I am convinced that their specimens refer to var. splendens, and not to var. phaeoleuca. Furrer-Ziogas (l.c.) also depicted clods of earth which is another indication of his misappliction. His statement that I. terrifera and I. phaeoleuca are extremely close and may probably not deserve specific rank, is correct.","Kuyper, Thomas W. 1986. A revision of the genus Inocybe in Europe : I. subgenus inosperma and the smooth-spored species of subgenus Inocybe, Rijksherbarium, Leiden, 247 p.; p. 216-217",,
  squalida C. Kaufholtz-Couture nom. prov.,Goniospora,Subcylindrical | Subamygdaliform | Laceroid | Rectangle | Subrectangle,Conical | Subconical,[9.5-15.5],[5.8-7.1],[1.6-2.6],Angular | Nodulose,Claviform | Cylindrical,[31-35],[9.5-11],[4-4],Yes,Fusiform | Metuloid,[35.6-53.6],[15.6-21.3],Yes | Sparse,[1-4],n/a,Yes,Ellipsoid | Ovoid | Pyriform | Subcylindrical,[14-25],[6.1-11.4],Yes,Fusiform | Metuloid,[55.6-59.6],[16.1-22],Yes,[2-3.7],Ammonioca -,Yes,Fusiform | Cylindrical | Claviform | Metuloid,[38.5-45.6],[12-16.9],Yes,n/a,n/a,No,n/a,n/a,n/a,No,n/a,n/a,n/a,PDAB -,Sandy soil,Beech | Birch | Oak,[8-15],[8-15],Subbulbous,n/a,n/a,Spermatic,Clypeus,,,,"Correspondence with B. Matheny and Thomas Kuyper. Both find similarities with Inocybe curvipes by microscopy, but the macromorphology is very different.","Kaufholtz-Couture, C. Taxonomy, macromorphological and microscopic descriptions, Fungiquebec, 2018; Collections examined : cKc0556; cKc0624; cKc0638; cKc0641",,,,
  squamata J.E. Lange,Leiospora,Subphaseoliform,Obtuse,[8.5-11],[5.5-7.5],[1.3-1.7],Smooth,Claviform,[30-37],[11-14],[2-4],Yes,Cylindrical | Claviform | Subutriform,[30-66],[10-20],Yes,n/a,n/a,n/a,n/a,n/a,n/a,No,n/a,n/a,n/a,No,n/a,n/a,Yes,Cylindrical | Claviform | Subutriform,n/a,n/a,No,n/a,n/a,No,n/a,n/a,n/a,No,n/a,n/a,n/a,n/a,Moist | Ground,Frondose | Poplar | Willow | Alder,[15-49],[15-49],Equal,n/a,n/a,Acidulous,Inosperma,Rimosae,,,"Occasionally specimens with an almost smooth pileus are met. The hyphae in the centre of the pileipellis do not or hardly sclerify in these specimens, and for that reason the pileus does not break into apressed scales. This character seems to be only a phenotypic response to moisture. In one collection, viz. Kuyper 1662, both character variants were found. Specimens growing in sheltered conditions were relatively smooth, but specimens growing in exposed situation were distinctly appressedly scaly. No taxonomic status can therefore be accorded to those smooth variants. 2) Inocybe squamata, especialy its variant with a rather smooth pileus, cannot always easily be separated from the extremely variable I. rimosa (Bull.: Fr.) Kumm. The latter species, however, has a more prominent umbo, a more strongly radially rimose pileal covering, a different smell and somewhat larger spores. I did not yet meet trully intermediate collections, and for that reason I. squamata is considered an autonomous species.","Kuyper, Thomas W. 1986. A revision of the genus Inocybe in Europe : I. subgenus inosperma and the smooth-spored species of subgenus Inocybe, Rijksherbarium, Leiden, 247 p.; p. 59-60",,,,
  squamosa Bres.,Leiospora,Amygdaliform,Conical,[8.5-10],[6.5-7.5],[1.2-1.5],Smooth,Claviform,[27-36],[9-12],[4-4],Yes,Cylindrical | Fusiform | Metuloid,n/a,n/a,Yes,n/a,n/a,n/a,n/a,n/a,n/a,Yes,Cylindrical | Fusiform | Metuloid,[64-78],[14-21],Yes,[2-2.5],n/a,Yes,n/a,n/a,n/a,n/a,n/a,n/a,Yes,n/a,n/a,n/a,n/a,n/a,n/a,n/a,n/a,Terrestrial,n/a,[10-15],[10-15],Subequal,n/a,n/a,Not distinctive,Inocybe,,,,Closely related to I. muricellata Bres. from which it maintly differs in remarkably broad spores. More material of I. squamosa seems necessary to assess its relationship to mediterranean variants of I. muricellata. The macroscopical description has been copied from Bresadola (l.c.).,"Kuyper, Thomas W. 1986. A revision of the genus Inocybe in Europe : I. subgenus inosperma and the smooth-spored species of subgenus Inocybe, Rijksherbarium, Leiden, 247 p.; p. 201-202",,,,
  
negative.code: microteroxantha Grund & D.E. Stuntz,Leiospora,Inequilaterally | Amygdaliform | Flat suprahilar dorsal area | Slight suprahilar depression,Conical | Subconical,[9-11],[4.5-6],n/a,Smooth,Claviform,[23-30],[8-11],[4-4],Yes,Versiform | Ventricose | Metuloid,n/a,n/a,Yes | No,n/a,n/a,Yes,Claviform,[24-33],[11-15],Yes,Ventricose | Metuloid,[38-68],[13-24],Yes | No,[3-5.5],n/a,Yes,Versiform | Ventricose | Metuloid,[50-72],[16-22],Yes | No,n/a,n/a,Yes,Versiform | Ventricose | Metuloid,[51-80],[11-20],Yes | No,n/a,n/a,n/a,n/a,Soil,Hardwoods | Conifers,[10-17],[10-17],Not bulbous,n/a,n/a,Spermatic | Not distinctive,,,,,"Our collections might be taken for depauperate specimens of Inocybe ochraceomarginata, but lack dermatocaulocystidia at the base of the stipe. We have been unable to find a record of an Inocybe with the particular combination of features of our species, i.e., bicolorous pileus (tawny and yellow), entirely pruinose stipe, smooth spores, and thick-walled pleurocystidia. Older specimens of I. langei Heim (Heim, R. 1931. Le genre Inocybe. Encyclop. Mycol. I. Lechevalier & Fils, Paris. 429 p.) in which the pure white stipe has become pallid yellow resemble our species, but have much smaller spores (7-8 x 4-5 micron), and pleurocystidia that do not exceed 50 micron long.","Grund, D.W.; Stuntz, D.E. Nova Scotian Inocybes VI., Mycologia, vol. 73, 1981; p. 658-659",,,,
  mimica Massee,Leiospora,Indistinctly phaseoliform,Obtuse | Subconical,[12-18.5],[6-9],[1.8-2.5],Smooth,Claviform,[29-39],[9-12],[4-4],Yes,Cylindrical | Claviform,[44-60],[14-19],No,n/a,n/a,n/a,n/a,n/a,n/a,No,n/a,n/a,n/a,No,n/a,n/a,n/a,n/a,n/a,n/a,n/a,n/a,n/a,n/a,n/a,n/a,n/a,n/a,n/a,n/a,n/a,n/a,Soil,Frondose,[50-65],[50-65],Not bulbous,n/a,n/a,Not distinctive,Inosperma,Rimosae,,,The macroscopical description is a translation of the description as given by Romagnesi (in Beih. Sydowia 8 : 349. 1979).,"Kuyper, Thomas W. 1986. A revision of the genus Inocybe in Europe : I. subgenus inosperma and the smooth-spored species of subgenus Inocybe, Rijksherbarium, Leiden, 247 p.; p. 58",,,,
  minima Peck,Leiospora,Inequilaterally | Amygdaliform,Conical | Papilla,[7-10],[4.5-6],n/a,Smooth,Claviform,[16-30],[6.5-13],[4-4],Yes,Ventricose | Metuloid,[44-70],[18-26],Yes | No,n/a,n/a,Yes,Sphaeropedunculate,[38-40],[11-24],Yes,Ventricose | Metuloid,[44-72],[9-24],Yes | No,n/a,n/a,Yes,Fusiform | Ovoid | Metuloid,[33-55],[11-18],No,n/a,n/a,No,n/a,n/a,n/a,No,n/a,n/a,n/a,n/a,Ground | Moss,Maple | Spruce,[6-14],[6-14],Not bulbous,n/a,n/a,Spermatic,,,,,"Inocybe minima belongs to series Lacerae (Fr.) Sacc. [Type species, I. lacera (Fr.) Kummer] and to the stirps Minima. The members of this stirps are small fungi with uniformly brown pilei, solid, pallid stipes, and thick-walled pleurocystidia. Other Inocybes with slender stems and brown pilei do not have all the above characters. Inocybe microspora J. Lange is the only other species that shares with I. minima the dark red-brown, gray-brown, or umber pileal coloration along with the dingy pallid stipe. Inocybe microspora has definitely smaller spores (6.5-7 micron long), shorter pleurocystidia (rarely exceeding 60 micron), and a longer stipe (usually 30-40 mm long) that becomes flushed with brown in age. Inocybe infelix Pk. (probably in the I. lacera Fr. complex) resembles I. minima, but has quite different spores and cystidia. Inocybe minima is unique because of its small size. This Inocybe has been reported twice from Canada by Smith and Wehmeyer (Stunyz, D. E. 1954. Studies of the genus Inocybe II. New and noteworthy species from Michigan. Pap. Michigan Acad. Sci. 39 : 53-84), by Wehmeyer (9) and by Bisby et al. (Bisby, G. R., et al. 1938. The fungi of Manitoba and Saskatchewan. Nat. Res. Council, Ottawa, Canada. 185 p., 8 pls). All three reports were unaccompanied by any descriptive data.","Grund, D.W.; Stuntz, D.E. Nova Scotian Inocybes VIII., Mycologia, vol. 76(4), 1984; p. 733-735",,,,
  mixtilis (Britzelm.) Sacc.,Goniospora,Subpolygonale,Conical | Subconical,[8-10],[5.5-7],n/a,9 to 10 rather prominent nodules,Claviform,[22-25],[7-12],[4-4],Yes,Ventricose | Metuloid,[38-50],[21-27],Yes | No,[2.5-3],n/a,Yes,Claviform,[16.5-23],[8-9],Yes,Ventricose | Metuloid,[40-55],[15-24],Yes,[5.5-8],n/a,Yes,Ventricose | Claviform | Metuloid,[35-66],[16-29],Yes,[2.5-3],n/a,Yes,Ventricose | Claviform | Metuloid,n/a,n/a,Yes,[2.5-3],n/a,n/a,n/a,Ground,Hemlock | Hardwoods,[15-35],[15-35],Marginate bulbous,n/a,cookei,Spermatic,Clypeus,Marginatae,Praetervisae,,"This Inocybe, one of the commonest species on the Pacific coast of North America, was reported in 1947 from Washington (Stuntz, D. E. 1947. Studies in the Genus Inocybe. I. New and noteworthy species from Washington. Mycologia 39 : 21-55; p. 48) as I. trechispora (Berk.) Karst., following the concept of that species by C. H. Kauffman. The salient features of the species are the silky-lubricous pileus in various combinations of fulvous and brassy yellow, the medium-sized, prominently nodulose spores, the ventricose pleurocystidia with thick walls, and, above all, the pale yellow, conspicuously flat-marginate bulb of the stipe. By these criteria, there is no doubt that our specimens represent I. trechispora sensu Kauffman, and I. mixtilis sensu Kuehner, and that those two concepts apply to one and the same taxon. Kuehner, while admitting the identity of his fungus with I. trechispora sensu Kauffman and Bresadola, chose, because of differing ""interpretations"" of Berkeley's species, to abandon that specific epithet, and use instead Britzlmayr's name. The matter could have been settled by examining Berkeley's type, but that was not done until relatively recently. Pegler and Young (Pegler, D. N., and Young, T. W. K. 1972. Basidiospore form in the British species of Inocybe. Kew Bull. 26 : 499-538) in studying spores of British Inocybes, examined the long-neglected types of several species, including Agaricus (Inocybe) trechisporous Berk. Their results make it clear that I. trechispora and I. mixtilis sensu Kuehner (possibly no one ever will know what I. mixtilis sensu Britzlmayr really is) are not identical. Inocybe trechispora does not have a marginate bulb and is excluded from the section Martinatae which includes I. mixtilis. There are other less important differences such as slight differences in the spores and paler color of I. trechispora, but the question of the relation of these two taxa now can be considered settled.","Grund, D.W.; Stuntz, D.E. Nova Scotian Inocybes V., Mycologia, vol. 72, 1980; p. 679-680",,,,
  monochroa J. Favre,Leiospora,Subamygdaliform,Conical,[9.5-11],[5.5-6.5],[1.7-1.9],Smooth,Claviform,[33-41],[9-12],[4-4],Yes,Cylindrical | Subfusiform | Sublageniform | Metuloid,n/a,n/a,Yes,n/a,n/a,Yes,Claviform,n/a,n/a,Yes,Cylindrical | Subfusiform | Sublageniform | Metuloid,[61-94],[11-17],Yes,[1.5-2],n/a,Yes,Cylindrical | Subfusiform | Sublageniform | Metuloid,n/a,n/a,n/a,n/a,n/a,No,Caulocystidioid hairs,n/a,n/a,No,n/a,n/a,n/a,n/a,Calcareous soil,Dryas octopetala | Willow | Pine,[10-24],[10-24],Subequal,n/a,n/a,Acidulous | Spermatic,Inocybe,,,,Inocybe monochroa comes very close to I. albovelutipes Stangl but differs in having larger spores and more slender cystidia with a thicker wall. More material is needed for a better evaluation of the relationships between both species.,"Kuyper, Thomas W. 1986. A revision of the genus Inocybe in Europe : I. subgenus inosperma and the smooth-spored species of subgenus Inocybe, Rijksherbarium, Leiden, 247 p.; p. 119-120",,,,
  "monticola Kropp, Matheny et Nanagyulyan sp. nov.",Leiospora,Amygdaliform,Subconical,[8-10.5],[5-6.5],n/a,Smooth,Claviform,[25-34],[8-11],[4-4],Yes,Fusiform | Ventricose | Metuloid,[48-70],[13-18],Yes,[1-2.5],n/a,n/a,n/a,n/a,n/a,Yes,Fusiform | Ventricose | Metuloid,[53-75],[13-20],Yes,[1-2.5],n/a,Yes,n/a,n/a,n/a,Yes,n/a,n/a,Yes,Irregular,n/a,n/a,Yes,n/a,n/a,n/a,n/a,Soil in montane environments,Trembling aspen,[13-60],[13-60],Not bulbous,n/a,n/a,Spermatic | Raphanoid,,,,,"Even though the collections of Inocybe monticola were made in late May or June at the relatively high elevations represented, this is early in the growing season and can be considered vernal. Inocybe monticola thus far has been found only in montane habitats at intermediate elevations in the western United States. How common or widespread this taxon is remains unknown. It has not been seen fruiting again at the type locale in the 10 y since it was first collected there. For the time being we can say only that it occurs in Oregon and Utah and that it does not appear to be part of the mycoflora that is seen annually. I. monticola material originally was identified as I. splendens var splendens sensu Kuyper (1986) who synonymized I. splendens Heim, I. alluvionis Stangl & Veselsky, and I. terrifera Kuehner. Inocybe monticola fits the concept for I. splendens sensu lato reasonably well, but based on our sequence data it differs phylogenetically from European material representing the species. The European material that we used in our phylogenetic work best fits I. splendens sensu Heim (1931) that is characterized by dark pileus colors and a distinct basal bulb on the stipe. Inocybe monticola differs from Heim's concept of I. splendens by lacking a distinct basal bulb on the stipe. Inocybe alluvionis is characterized by a lighter pileus color than I. monticola and by having a submarginate basal bulb on the stipe that is not present on I. monticola. Inocybe terrifera lacks a basal bulb but can be separated from I. monticola by its lighter pileus colors. Inocybe monticola comes closest to I. phaeoleuca Kuehner in having dark pileus colors and lacking a basal bulb. However its DNA sequences differ sharply from that of the European exemplar of I. phaeoleuca Kuehner and its stature is considerably more robust than that of I. phaeoleuca. In addition a velipellis is present on most of the material of I. monticola and this is usually absent from I. phaeoleuca. One rare European species, I. glabrescens Velen, also has some similarity to I. monticola but differs by having a rimose pileus margin and a more slender stipe that discolors yellowish in age. The material on which I. monticola is based does not belong to either I. splendens sensu Heim or I. phaeoleuca sensu Kuehner, and as mentioned above there are morphological reasons to exclude it from both I. alluvionis and I. terrifera.","Kropp, Bradley R.; Matheny, P. Brandon,; Nanagyulyan, Siranush G. Phylogenetic taxonomy of the Inocybe splendens group and evolution of supersection ""Marginatae"", Mycologia, 102(3), 2010, pp. 560-573 ; p.569-570",,,,
  mucidiolens (Grund & D.E. Stuntz) Matheny,Leiospora,Elongate | Subreniform,Obtuse,[10-12],[5-6],n/a,Smooth,Claviform,[33-42],[8-10],[4-4],Yes,Claviform | Subcylindrical,[33-55],[10-22],No,n/a,n/a,n/a,n/a,n/a,n/a,No,n/a,n/a,n/a,No,n/a,n/a,Yes,Claviform,[27-50],[10-22],No,n/a,n/a,No,n/a,n/a,n/a,No,n/a,n/a,n/a,n/a,Moss | Needle,Conifers,[10-40],[10-40],Somewhat bulbous,n/a,n/a,Green-corn,,,,,"In its strong green-corn odor var. mucidiolens differs markedly from I. calamistrata var. calamistrata, which always smells like a mixture of fresh raw fish and pine resin. The two taxa are similar in other respects, both showing a certain amount of variability in such features as size, amount of green color in the stipe, and reddening of exposed context.","Grund, D.W.; Stuntz, D.E. Nova Scotian Inocybes II., Mycologia, vol. 62, 1970; p. 929-931","A comparison between the protologues of Agarigus calamistratus Fr.: Fr. and A. hirsutus Lasch: Fr. suggests that both names refer to the same taxon; representative material from various parts of Europe did not give any reason to assume the existence of two different species or infraspecific taxa. Considering the variability in the smell of various collections of I. calamistrata, it seems likely that var. mucidiolens Grund & Stuntz in Mycologia 62 : 929 (1970) is only a minor variant without autonomous status. I did not study the type, however.","Kuyper, Thomas W. 1986. A revision of the genus Inocybe in Europe : I. subgenus inosperma and the smooth-spored species of subgenus Inocybe, Rijksherbarium, Leiden, 247 p.",,Brandon Matheny
  multicoronata A.H. Sm.,Goniospora,Globose | Ellipsoidal,Conical,[12-14.5],[10-12],n/a,"3 to 6 prominent nodules, blunt or truncate warts",Claviform,[38-44],[13-17],[4-4],Yes,Subventricose | Ventricose | Metuloid,n/a,n/a,Yes | No,n/a,n/a,n/a,n/a,n/a,n/a,Yes,Subventricose | Ventricose | Metuloid,[55-100],[13-24],Yes | Sparse | No,[3-4.5],n/a,Yes,Lageniform | Claviform,n/a,n/a,Yes | Sparse | No,n/a,n/a,Yes,Fusiform,n/a,n/a,Yes | Sparse | No,n/a,n/a,n/a,n/a,Moss | Needles,Spruce,[5-20],[5-20],Equal,n/a,n/a,n/a,Clypeus,,,,"This Inocybe apparently has not been found outside of Nova Scotia, and has not been reported since it was described in 1939. Its conical scaly pileus, slender, entirely fibrillo-pruinose stipe, and especially the spores with multicoronate warts, make it a very distinctive species. Only two other Inocybes with similar spores have been described, I. lasseri Dennis from Venezuela, and I. petchii Boedijn from Ceylon and Java. These have some features other than the spores in common with I. multicoronata, which thus is the boreal representative of a stirps with world-wide distribution.","Grund, D.W.; Stuntz, D.E. Nova Scotian Inocybes I., Mycologia, vol. 60, 1968; p. 418-420",,,,
  muricellata Bres.,Leiospora,Subamygdaliform,Conical,[8-13],[5-7],[1.5-2.1],Smooth,Claviform,[24-32],[8-11],[2-4],Yes,Cylindrical | Fusiform | Sublageniform | Lageniform | Metuloid,n/a,n/a,Yes,n/a,n/a,Yes,Pyriform | Claviform,n/a,n/a,Yes,Cylindrical | Fusiform | Sublageniform | Lageniform | Metuloid,[54-95],[10-22],Yes,[2-3],n/a,Yes,Cylindrical | Fusiform | Sublageniform | Lageniform | Metuloid,n/a,n/a,n/a,n/a,n/a,Yes,n/a,n/a,n/a,n/a,n/a,n/a,Yes,n/a,Calcareous soil,Spruce | Pine | Cedar | Beech | Oak | Hazel | Blue-beech | Birch | Alder,[7-38],[7-38],Equal | Claviform | Subbulbous,n/a,n/a,Nihil | Bitter almonds | Subspermatic,Inocybe,,,,"1) Inocybe muricellata is a rather variable species both in macroscopical and microscopical respects, and it has seemed possible to subdivide this material into two taxa maintly differing in colour of stipe correlated with some microscopical characters. Collections which show a distinctly purplish red or lilac red apex of stipe generally possess smaller spores and cystidia. However, when more material was studied these microscopical characters were found to show too much overlap to be reliable for a formal subdivision. Specimens with only a faint pinkish or orange tinge at the stipe apex were also encountered, indicating that this character is more gradual too. Both variants do not seem to differ significantly in their ecology, although the variant with the reddish stipe seems to prefer somewhat more natural vegetation, but this character also proved insufficient for a formal subdivision. The macroscopical structure of the pileipellis is dependent on the age of the specimens and the development of the velipellis. These phenotypical differences between strongly squarrose specimens and tomentose specimens are devoid of taxonomical importance. 2) In the collection that I made myself I never noted a smell of bitter almonds. It might therefore be possible that the mention of this smell in several collections is based on biased observations. This might be true for instance in the description by J. Lange (Fl. agar. dan. 3: 78. 1938) where a faint smell of bitter almonds is noted, although the description definitely refers to I. muricellata. 3) Inocybe hirtella Bres. differs in having a conspicuous smell of bitter almonds, being somewhat brighter yellow, and possessing smaller, slenderly clavate cystidia with less bright yellow walls. It never shows such conspicuous reddish tinges at the stipe apex.","Kuyper, Thomas W. 1986. A revision of the genus Inocybe in Europe : I. subgenus inosperma and the smooth-spored species of subgenus Inocybe, Rijksherbarium, Leiden, 247 p.; p. 199-201",,,,
  mycenoides Kuyper,Leiospora,Subamygdaliform,Subconical,[8-11],[5.5-6.5],[1.5-1.8],Smooth,Claviform,[23-29],[7-9],[4-4],Yes,Cylindrical | Claviform | Subfusiform | Metuloid,n/a,n/a,Yes,n/a,n/a,Yes,Claviform,n/a,n/a,Yes,Cylindrical | Claviform | Subfusiform | Metuloid,[31-50],[10-16],Yes,[1.5-2.5],n/a,Yes,Cylindrical | Claviform | Subfusiform | Metuloid,n/a,n/a,n/a,n/a,n/a,Yes,n/a,n/a,n/a,n/a,n/a,n/a,Yes,n/a,Soil,Beech | Basswood | Larch,[6-30],[6-30],Equal,n/a,n/a,Subspermatic,Inocybe,,,,"Recognisable because of its mycenoid habit, completely pruinose stipe and short, cylindrical to subfusiform cystidia. The collection from the Netherlands consisted unfortunately of one specimen only of which the lamellae were less yellow than Metrod described for his I. citrinofolia.","Kuyper, Thomas W. 1986. A revision of the genus Inocybe in Europe : I. subgenus inosperma and the smooth-spored species of subgenus Inocybe, Rijksherbarium, Leiden, 247 p.; p. 210-212",,,,
  