; (B) exploration of genetic manipulation of plant pests or noxious weeds; (C) enhancement of pest-resistance mechanisms of hosts; and (D) development of other strategies for restoring individual tree species; and (2) develop, and disseminate to the public, tools and information based on the research conducted under paragraph (1). (c) Applications An eligible entity seeking to receive a grant under this section shall submit to the Secretary an application at such time, in such manner, and containing such information as the Secretary may require, including a description of a comprehensive forest restoration research program to be carried out by the eligible entity using the funds received through the grant. (d) Use of funds (1) In general An eligible entity receiving a grant under this section shall use the funds received through the grant to conduct research intended to address specific questions relating to the recovery of tree species that are native to the United States and suffering severe levels of mortality caused by nonnative plant pests or noxious weeds. (2) Matching requirement (A) In general An eligible entity receiving a grant under this section shall provide matching funds from non-Federal sources in an amount equal to not less than 20 percent of the grant. (B) Indirect costs (i) In general Indirect costs charged against a grant awarded under this section shall not exceed 30 percent of the total Federal funds provided under the grant award. (ii) Inclusions Indirect costs described in clause (i) shall include— (I) equipment used in relation to the grant; (II) capital improvements of facilities that are necessary to carry out the grant; (III) accounting costs, personnel costs, and administrative costs incurred by an eligible entity necessary to carry out the grant