; (B) a State college, university, or agency; (C) a State-licensed veterinarian; (D) a wildlife sanctuary that— (i) cares for prohibited wildlife species; (ii) is a corporation that is— (I) exempt from taxation under section 501(a) of the Internal Revenue Code of 1986; and (II) described in sections 501(c)(3) and 170(b)(1)(A)(vi) of that Code; (iii) does not commercially trade in any prohibited wildlife species, including offspring, parts, and byproducts of prohibited wildlife species; (iv) does not breed any prohibited wildlife species; (v) does not allow direct contact between the public and any prohibited wildlife species; and (vi) does not allow the transportation and display of any prohibited wildlife species off-site; (E) an entity that has custody of any prohibited wildlife species solely for the purpose of expeditiously transporting the prohibited wildlife species to a person described in this paragraph with respect to the prohibited wildlife species; or (F) an entity or individual that— (i) is in possession of any prohibited wildlife species that was born before the date of enactment of the Big Cat Public Safety Act ; (ii) not later than 180 days after the date of enactment of that Act, registers each individual animal of each prohibited wildlife species possessed by the entity or individual with the United States Fish and Wildlife Service; (iii) does not breed, acquire, or sell any prohibited wildlife species after the date of enactment of that Act; and (iv) does not allow direct contact between the public and any prohibited wildlife species. . 4. Penalties (a) Civil penalties Section 4(a)(1) of the Lacey Act Amendments of 1981 ( 16 U.S.C. 3373(a)(1) ) is amended— (1) by inserting (e), after (d), ; and (2) by inserting , (e), after subsection (d) . (b) Criminal penalties Section 4(d) of the Lacey Act Amendments of 1981 ( 16 U.S.C. 3373(d) ) is amended— (1) in paragraph (1)— (A) in subparagraph (A), by inserting (e), after (d), ; and (B) in subparagraph (B), by inserting (e), after (d),