Rubus cissoides
Introduction '''''Rubus cissoides''''', commonly called '''bush lawyer''' or '''tātarāmoa''' in te reo Māori, is a species of flowering plant in the family Rosaceae, endemic to New Zealand. Alan Cunningham described ''R. cissoides'' in 1839.Plants of this species of are perennial scrambling vines with compound leaves with 3-5 leaflets each up to 15 cm long, reddish prickles on the branches, white flowers from September to November and red berries from December to April. The conservation status of ''R. cissoides'' is Not Threatened, it is widespread on all three main islands of mainland New Zealand, and it has been used by Māori as food, medicines and construction materials.
Taxonomy and Etymology ''Rubus cissoides'' A.Cunn. is in the family Rosaceae. The species was described in 1839 by Alan Cunningham, based on the type specimen collected in 1826 by Cunningham from "dense forests of Whangaroa". The type specimen is lodged at Kew Herbarium (K000762091). The epithet ''cissoides'' means "ivy like" and is derived from the Greek words ''kissos'' (ivy, ''Cissus'') and ''-oides'' (likeness). ''Rubus cissoides'' is the same genus, ''Rubus,'' as the blackberry and raspberry. In addition to ''R. cissoides,'' there are four other endemic species of ''Rubus'' in New Zealand, including ''R. australis, R. parvus, R. schmideloides,'' and ''R. squarrosus.'' These can be distinguished from one another using prickle, leaf, inflorescence and fruit characteristics. ''Rubus cissoides'' is the most common native species in New Zealand, and has sharply toothed leaflets that are long and narrow. Plants of this species can grow very high - over 15 metres above the ground - and stems may be 17 cm in diameter.Many of the native New Zealand ''Rubus'' species, including ''R. cissoides,'' are commonly called bush lawyer or tātarāmoa in te reo Māori, and other vernacular names for this and other similar ''Rubus'' species in New Zealand include taramoa, akatātarāmoa, taraheke, and tātaraheke. They are considered to be "hook climbers", and are woody, low to high climbing vines or lianes that have backwardly curving hooked prickles, which allow the plant to grip, climb up and find support on shrubs or other vegetation.Close up of white flowers of ''Rubus cissoides''
Description ''Rubus cissoides'' has thin branches, serrated leaves and white flowers ''Rubus cissoides'' plants are dioecious vines up to 10 m long that scramble or climb on other vegetation. Young stems are covered in hooked reddish prickles, especially on the underside, whereas mature stems are without prickles and are hairless. Leaves are palmately or ternately compound, smooth and hairless, with up to 5 long and narrow leaflets (young plants) or 3–5 leaflets (mature plants). Petioles and petiolules have hooked prickles. Leaflets are variable in size and shape, 6–15 cm long and up to 6 cm wide, lanceolate or oblong, sharply toothed on the edges, with a pointed tip. Inflorescences are large and much-branched panicles of white flowers up to 20 cm long. Flowers have 5 sepals and 5 petals, and are either male (with numerous stamens, but with no style or only a rudimentary one) or female (with functional style, but with no stamens or only a few rudimentary stamens). The berries are around a centimetre in diameter, yellow to red, and made up of about 10–15 drupelets.
Distribution and habitat ''Rubus cissoides'' is endemic to and widespread in New Zealand, and occurs on all three of the main islands: North Island, South Island and Stewart Island. It is found in lowland and montane habitats, often in forests, but also in scrub and wetland margins, often in moist, sunny environments.
Conservation status ''Rubus cissoides'' is considered to be Not Threatened in the most recent assessment (2017–2018) under the New Zealand Threat Classification system for plants.
Life cycle and phenology In a seed germination experiment, 82% of ''Rubus cissoides'' seeds germinated, with some germinating in spring, remaining seeds germinating at a slow, steady rate over two years.Young plants spread over the forest floor until finding an appropriate shrub or vegetation for support. They can support themselves up to a height of about 60 cm before requiring support in the form of other vegetation. ''R. cissoides'' is insect-pollinated, and it flowers from September to November and fruits from December to April.
Herbivory and diseases The non-native herbivores, deer and possums, eat small amounts of the foliage or leaf litter of ''R. cissoides,'' and possums have been known to eat the berries and flowers of ''R. cissoides''.The blackberry rust fungus ''Phragmidium violaceum'', appears as black patches on the leaves and can cause mild infections in ''R. cissoides.''
Uses ''Rubus cissoides'' and other native New Zealand ''Rubus'' species were used by Māori and continue to have multiple uses, including as food, construction materials, or medicines. The fruits are eaten by birds and people, the branches can be used for making traps, and the crushed berries form a dye which is blue or purple in colour. Some of the ways Māori used ''Rubus'' plants medicinally include using the bark of the stem to treat abdominal pains, using root bark to treat diarrhea, and preparing crushed leaves to relieve chest congestions and colds.
Gallery File:Rubus cissoides A.Cunn. (AM AK376537).jpgFile:Rubus cissoides - BushLawyer.jpgFile:Rubus cissoides Lake Matheson Walk MRD 09.jpgFile:Rubus cissoides 239955101.jpgFile:Rubus cissoides 199185439.jpg
References
External links * * ''Rubus cissoides'' occurrence data from Australasian Virtual Herbarium*