Do branch circuits sharing conduit with sub-panel feeder wires require separate grounding?
No, one ground wire of sufficient size will do the trick

All the branch circuits in a single cable or conduit can share a single ground wire (and are often forced to when metal conduit is used as a ground path or when a multi-circuit cable is used).  This is permitted as a consequence of NEC 250.122(C):


  (C) Multiple Circuits. Where a single equipment grounding
  conductor is run with multiple circuits in the same raceway,
  cable, or cable tray, it shall be sized for the largest overcurrent
  device protecting conductors in the raceway, cable, or cable
  tray. Equipment grounding conductors installed in cable trays
  shall meet the minimum requirements of 392.10(B)(1)(c).