Bootstrapping (symmetry breaking) in evolution
I found a paper which gives four conditions that have to be met:


Condition 1: A rule should be enforced that individuals
must meet some minimal criterion (MC) before they
can reproduce, and that criterion must be nontrivial
Condition 2: The evolution of new individuals should
create novel opportunities for satisfying the MC
Condition 3: Decisions about how and where individuals
interact with the world should be made by the individuals
themselves
Condition 4: The potential size and complexity of
the individuals’ phenotypes should be (in principle) unbounded


Source: Soros, Stanley (2014): Identifying Necessary Conditions for Open-Ended Evolution through the Artificial Life World of Chromaria

Chromaria is "an experimental platform that allows the exploration of evolutionary theories in an intuitive and visually engaging way" from EPLEX, the Evolutionary Complexity Research Group at the University of Central Florida (UCF)