Is the information in the brain stored in the connections rather than neurons?
Is the information in the Brain stored in the connections rather than
  in the neurons ?


It depends on what information you are referring to.

The brain does not just store one type of information,the model of a grandmother cell and the model of an interconnected neuron network are 2 subsystems of a greater system working together perhaps in a competitive or complementary arrangement. The purpose of this system is to extract information from the environment and identify it again for the organisms benefit. 1.

So let's say we see something that will benefit us in the future or that we might want to avoid. This is a simplification, but as you see that something, neurons in your neocortex light up specifically in region V1 in the back of your head, they arrive there after a lot of computations along the way from your retina, but also project to other parts of your brain where they are perhaps processed consciously,they fire in an orderly fashion, in a map, this bouncing around of electrical signals represents your conscious, and unconscious processing of the image, so the information exists as a network of electrical signals. 2.

During recollection, ( there are different types of memories so the structure in charge could be the neocortex,basal ganglia, hippocampus or even a muscle)3. this same map ( or a smaller subset ) gets replayed. The mechanism(s) by which this information is first encoded and later retrieved is still not entirely understood, but it involves a map and a getaway or switchboard.4. This switchboard ties together different percepts ( a smaller network of electrical signals), so in a way the information is represented at the switchboard level, if you disable this switchboard, you can't make new memories or lose previous ones.

A grandmother cell in the scheme of what I just explained, is the result of another subsystem that helps recognize something. Once you successfully encode a percept ( thing we want to remember seeing) you might want to associate it with something ( good,bad,delicious,etc), at the top of a series of associations lies a grandmother cell, which in turn is a handy evolutionary way of signaling back to other systems when some previous information has been experienced. So in this way the information of whether a subset of relationships in between networks of electrical signals has been experienced and encoded lies here.

References/Sources:


The idea of grandmother cells and object recognition in general is summarized in the Object Recognition chapter of ( Cognitive Neuroscience,Gazzinga-3rd ed 222-225) along with the problems a grandmother cell introduces in object recognition (ie if you lose the grandmother cell you would lose all the information about your grandmother)
Koch( Quest for Consciousness-2004) chapters on vision provide a very readable account of how the information goes from retina to v1. Vision Science (Palmer) is a more thorough account. 
The chapter on learning and memory (Cognitive Neuroscience) gives an overview. 
(Gluck -2001-Gateway to memory) presents different models of this arrangement.