How exactly can an airship lose lift?
The reason is a temperature difference between the lift gas and the surrounding air, and probably water uptake by the hull when descending through clouds.

A given mass of hydrogen will create a constant lift force, regardless of pressure or altitude, when at equal pressure and temperature with the surrounding air. Therefore, a change in altitude will not change the lift a rigid airship creates. Ideally.

However, air gets colder the higher you climb. The sun heats the atmosphere from below, by way of heating the ground, and space cools it from the top. Therefore, on many days the temperature gradient is bigger than its adiabatic value - that is the way thermals work! L-55 stayed at altitudes where air is -32°C according to the Standard Atmosphere. When descending, the surrounding air got warmer and also warmed up the lift gas, but only slowly. This means that, depending on the rate of descent, the lift gas lagged behind in temperature relative to the air, and this temperature difference reduced its lift capacity.

Note that adiabatic heating will already heat a gas when being compressed. The lapse rate of the atmosphere must be above the dry adiabatic value for this mechanism to work, which it is on many days. Especially behind a cold front. Note that L-55 encountered strong winds - just what you find in and behind a cold front. So it is safe to conclude that L-55 flew in labile air, and when it descended, that motion became unstable, at least close to the ground.

Kapitän Flemming simply descended too quickly. Slowing things down would have warmed the lift gas more, and less lift would have been lost. But delaying the descent has dangers of its own: The gas bags back then were made from goldbeater skin and had a certain amount of seepage. To compensate, Zeppelins started their trips with several tons of ballast water onboard, which was progressively dropped during the many hours of a normal trip. Taking a detour over France delayed the trip, so he was running short on time.

Normally, a loss of lift can be compensated by dynamic lift. With some angle of attack, an airship can create up to 20% of its weight in dynamic lift - as long as all engines are running. L-55 was blown south during the night after attacking Hull and Birmingham, and found itself far more south than assumed when dawn allowed the crew to get a ground fix. When back over Germany, L-55 ran out of fuel and dynamic lift was no longer available to compensate for the lower lift gas temperature. It made a rough landing in the Thuringian countryside near Tiefenort and had to be written off.

Literature: Heinz Urban, Zeppeline der kaiserlichen Marine 1914 bis 1918