Can android collect info about wifi routers even if Google Play Service is disabled?
Android collects information about WiFi routers through the normal course of WiFi operation. You cannot get around this fact if you want to use WiFi. During normal WiFi operation the device listens for management packets like beacons that are sent from WiFi access points, and this information, which is used to communicate with WiFi APs, can be used to estimate location with an appropriate database (Google, Apple, Skyhook, etc.). 

In some older versions of Android, it was actually possible to read the AP information with network permissions only and estimate the location using this information with a custom query to a database (not directly through Google's service). In more recent versions (maybe starting at 6?), I think this loophole has been fixed, so that the results of a wireless scan no longer give AP information that can be used to compute location. See https://stackoverflow.com/q/34957118/4107809. 

I don't know anything about the default settings.