Flowers that are located directly in line with the sun are shown by waggle runs in an upward direction on the vertical combs . If the flowers are at an angle to the right or left of the sun , the waggle run is done at the same angle to the upward direction . Flowers located directly in line with the sun are represented by waggle runs in an upward direction on the vertical combs , and any angle to the right or left of the sun is coded by a corresponding angle to the right or left of the upward direction .