salmon runs
... that the salmon caught by grizzly bears during salmon runs contribute significantly to the nitrogen cycle in adjacent woodlands where bears urinate, defecate, and drop partially eaten fish carcasses?
... that the salmon caught by <a href="./Grizzly_bear" id="mwD94" rel="mw:WikiLink" title="Grizzly bear">grizzly bears</a> <i id="mwD98">(pictured)</i> during <b id="mwD-A"><a href="./Salmon_run" id="mwD-E" rel="mw:WikiLink" title="Salmon run">salmon runs</a></b> contribute significantly to the <a href="./Nitrogen_cycle" id="mwD-I" rel="mw:WikiLink" title="Nitrogen cycle">nitrogen cycle</a> in adjacent woodlands where bears urinate, defecate, and drop partially eaten fish carcasses?