theoretical ecology
... that mathematical models developed in theoretical ecology show that complex food webs tend to be unstable?
... that <a href="./Mathematical_model" id="mwC9E" rel="mw:WikiLink" title="Mathematical model">mathematical models</a> developed in <b id="mwC9I"><a href="./Theoretical_ecology" id="mwC9M" rel="mw:WikiLink" title="Theoretical ecology">theoretical ecology</a></b> show that complex <a href="./Food_chain" id="mwC9Q" rel="mw:WikiLink" title="Food chain">food webs</a> <i id="mwC9U">(pictured)</i> tend to be unstable?