ion ▁W ost l , ▁R iy anto , ▁Ham id y , ▁K urn ia wan , ▁Smith , ▁and ▁Har vey , ▁ 2 0 1 7 ▁Phil aut us ▁vent rim ac ul atus ▁W ost l , ▁R iy anto , ▁Ham id y , ▁K urn ia wan , ▁Smith , ▁and ▁Har vey , ▁ 2 0 1 7 ▁Gen us : ▁Pol yp ed ates ▁Tsch udi , ▁ 1 8 3 8 ▁Pol yp ed ates ▁col lett i ▁ ▁( B ou len ger , ▁ 1 8 9 0 ) ▁Pol yp ed ates ▁is k and ari ▁R iy anto , ▁M ump uni , ▁and ▁Mc Gu ire , ▁ 2 0 1 1 ▁Pol yp ed ates ▁le u com yst ax ▁ ▁( G ra ven hor st , ▁ 1 8 2 9 ) ▁Pol yp ed ates ▁mac rot is ▁ ▁( B ou len ger , ▁ 1 8 9 1 ) ▁Pol yp ed ates ▁mut us ▁ ▁( Smith , ▁ 1 9 4 0 ) ▁Pol yp ed ates ▁o til oph us ▁ ▁( B ou len ger , ▁ 1 8 9 3 ) ▁Pol yp ed ates ▁pseud ot il oph us ▁M ats ui , ▁Ham id y , ▁and ▁K ura ishi , ▁ 2 0 1 4 ▁Gen us : ▁Rh ac op hor us ▁Ku hl ▁and ▁Van ▁Hass elt , ▁ 1 8 2 2