▁have ▁been ▁described ▁as ▁some ▁of ▁the ▁an atomic ▁changes ▁associated ▁with ▁hyper h yd ric ity . ▁ ▁C aus es ▁The ▁main ▁causes ▁of ▁hyper h yd ric ity ▁in ▁plant ▁t issue ▁culture ▁are ▁those ▁factors ▁trigger ing ▁ox id ative ▁str esses ▁such ▁as ▁high ▁salt ▁concentration , ▁high ▁relative ▁hum id ity , ▁low ▁light ▁intensity , ▁gas ▁accum ulation ▁in ▁the ▁atmosphere ▁of ▁the ▁jar , ▁length ▁of ▁time ▁intervals ▁between ▁sub c ult ures ; ▁number ▁of ▁sub c ult ures , ▁concentration ▁and ▁type ▁of ▁g elling ▁agent , ▁the ▁type ▁of ▁expl ants ▁used , ▁the ▁concentr ations ▁of ▁micro element ▁and ▁h orm onal ▁im bal ances . ▁Hyper h yd ric ity ▁is ▁commonly ▁apparent ▁in ▁liquid ▁culture - g rown ▁plants ▁or ▁when ▁there ▁is ▁low ▁concentration ▁of ▁g elling ▁agent . ▁High ▁am mon ium ▁concentration ▁also ▁cont ributes ▁to ▁hyper h yd ric ity . ▁ ▁Control ▁ ▁Hyper h yd ric ity ▁can ▁be ▁monitor ed ▁by ▁modifying ▁the ▁atmosphere ▁of ▁the ▁culture ▁vessels . ▁Ad just ing ▁the ▁relative ▁hum id ity ▁in ▁the ▁vessel ▁is ▁one ▁of ▁the ▁most ▁important ▁parameters ▁to ▁be ▁controlled . ▁Use ▁of ▁gas - per me able ▁memb ran es ▁may ▁help ▁in ▁this ▁regard ▁as ▁this ▁allows ▁increased ▁exchange ▁of ▁water ▁v apor ▁and ▁other ▁g ases ▁such ▁as ▁eth yl ene ▁with ▁the ▁surrounding ▁environment . ▁Using ▁higher ▁concentration ▁of ▁a ▁g elling ▁agent , ▁on ▁top ▁of ▁the