▁( 1 9 9 9 ) ▁places ▁the ▁genus ▁H oul let ia ▁within ▁the ▁" Ac in eta - Ver wand t schaft sg ru ppe ," ▁al lied ▁with ▁Ac in eta , ▁Lu ed dem ann ia ▁and ▁Vas que ziel la ▁within ▁the ▁larger ▁Stan ho pe inae ▁Alliance . ▁ ▁Wh itten , ▁Williams , ▁and ▁Ch ase ▁distinguish ▁the ▁H oul let ia ▁cla de ▁among ▁the ▁Stan ho pe inae , ▁compr ising ▁Hor ich ia , ▁H oul let ia , ▁Pap hin ia , ▁Sch lim m ia , ▁and ▁Tre v oria . ▁They ▁write : ▁ ▁" As ▁presently ▁defined , ▁H oul let ia ▁consists ▁of ▁two ▁morph olog ically ▁distinct ▁groups . ▁The ▁group ▁containing ▁H . ▁bro ck el h urst iana ▁( the ▁type ▁species ), ▁H . ▁t igr ina , ▁H . ▁od or at iss ima ▁( L inden ▁ex ▁Lind l .), ▁and ▁H . ▁jur uen ensis ▁( H oe h ne ) ▁have ▁open , ▁res up inate ▁flowers ▁that ▁are ▁heavily ▁sp otted ▁in ▁red - b rown . ▁The ▁ep ich ile ▁is ▁tri angular ▁and ▁the ▁hy po ch ile ▁be ars ▁a ▁pair ▁of ▁cur ved , ▁ac ute ▁pro jections ; ▁the ▁lip ▁shares ▁many ▁features ▁of ▁the ▁lip ▁of ▁Pap hin ia . ▁The ▁vis cid ium ▁is ▁narrow , ▁approximately ▁the ▁same ▁width ▁of ▁the ▁long ▁st ipe , ▁and ▁the ▁pol lin aria ▁are