▁and ▁ch ir onom id ▁lar va e . ▁ ▁It ▁may ▁also ▁feed ▁on ▁he pt agen i id ▁n ym ph s ▁on ▁the ▁unders ide ▁of ▁rocks . ▁ ▁They ▁become ▁sex ually ▁m ature ▁when ▁they ▁are ▁greater ▁than ▁one ▁year ▁old ▁and ▁at ▁least ▁ ▁long . ▁Research ▁on ▁the ▁cit ico ▁d arter ▁( E the ost oma ▁sit iku ense ), ▁another ▁member ▁of ▁the ▁dus ky tail ▁complex , ▁has ▁shown ▁that ▁it ▁can ▁produce ▁five ▁to ▁seven ▁cl ut ches ▁per ▁year ▁with ▁cl utch ▁sizes ▁recorded ▁up ▁to ▁ 1 0 1 ▁eggs . ▁ ▁The ▁cit ico ▁d arter ▁spawn s ▁at ▁water ▁temper atures ▁between ▁. ▁ ▁The ▁t ux edo ▁d arter ▁is ▁known ▁to ▁spawn ▁under ne ath ▁sl ab ro cks ▁or ▁b ould ers . ▁ ▁The ▁t ux edo ▁d arter ▁pre fers ▁river ▁sections ▁that ▁are ▁ ▁wide ▁and ▁ ▁deep . ▁Common ▁substr ates ▁for ▁the ▁species ▁are ▁sl ab rock , ▁c ob ble , ▁and ▁b ould ers ▁where ▁they ▁build ▁n ests ▁up ▁to ▁ ▁up stream ▁which ▁contain ▁the ▁eggs ▁of ▁one ▁or ▁more ▁females ▁but ▁are ▁guard ed ▁by ▁a ▁single ▁male . ▁The ▁t ux edo ▁d arter ▁clusters ▁its ▁eggs . ▁ ▁Conserv ation ▁E . ▁per cn ur um ▁was ▁first ▁recorded ▁in ▁the ▁C umber land ▁River ▁in ▁ 1 9 7 6 . ▁ ▁The ▁four ▁separate ▁populations , ▁now ▁four ▁different ▁species ,