▁genus ▁name ▁Princi p ial a ▁was ▁co ined ▁by ▁the ▁research ers ▁as ▁a ▁combination ▁of ▁" Pr inci pi ", ▁in ▁honor ▁of ▁Italian ▁ent om ologist ▁Maria ▁Princi pi ▁and ▁the ▁Latin ▁word ▁al a ▁meaning ▁" wing ". ▁ ▁The ▁specific ▁ep ith et ▁" inc erta " ▁is ▁derived ▁from ▁the ▁Latin ▁inc ert us , ▁in ▁reference ▁to ▁the ▁uncertain ▁relationship ▁of ▁the ▁genus ▁within ▁I th on idae . ▁ ▁Two ▁years ▁later ▁a ▁group ▁of ▁research ers ▁led ▁by ▁pale o ent om ologist ▁James ▁Je pson ▁described ▁the ▁second ▁species ▁P . ▁rud g wick ensis . ▁ ▁The ▁team ▁chose ▁the ▁species ▁name ▁" r ud g wick ensis " ▁as ▁a ▁reference ▁to ▁the ▁Rud g wick ▁Br ick works ▁where ▁the ▁foss il ▁was ▁found . ▁ ▁Over all ▁the ▁ve in ▁structure ▁of ▁Princi p ial a ▁is ▁most ▁similar ▁to ▁the ▁genus ▁All or apis ma , ▁known ▁from ▁Y pres ian ▁foss ils ▁found ▁in ▁the ▁north western ▁United ▁States . ▁ ▁The ▁wings ▁of ▁both ▁gener a ▁show ▁the ▁ap ical ▁end ▁of ▁the ▁R s ▁ve in ▁f used ▁to ▁the ▁MA ▁ve in . ▁ ▁Both ▁have ▁similar ▁position ing ▁and ▁sh aping ▁of ▁the ▁MP ▁ve in ▁and ▁the ▁Cu A ▁ve in ▁running ▁parallel ▁to ▁the ▁MP ▁ve in . ▁ ▁As ▁such , ▁Mak ark in ▁and ▁Arch ib ald ▁suggested ▁the ▁two ▁be ▁grouped ▁in ▁an ▁inform al ▁sup rag ener