▁of ▁the ▁Tot ol apa ▁region ▁is ▁currently ▁identified ▁as ▁E oc ene ▁in ▁age , ▁but ▁the ▁fa una ▁of ▁the ▁am ber ▁is ▁very ▁similar ▁to ▁both ▁the ▁Sim oj ov el ▁fa una ▁and ▁to ▁Domin ican ▁am ber , ▁indicating ▁that ▁a ▁re ass ess ment ▁of ▁the ▁ge ology ▁may ▁be ▁needed . ▁ ▁The ▁hol ot ype ▁was ▁first ▁studied ▁by ▁pale o ent om olog ists ▁David ▁C oty , ▁Rom ain ▁G arr ou ste ▁and ▁André ▁Nel , ▁of ▁the ▁Mus é um ▁National . ▁ ▁Their ▁type ▁description ▁of ▁the ▁species ▁was ▁published ▁in ▁the ▁Ann ales ▁de ▁la ▁Société ▁Ent om ologique ▁de ▁France ▁in ▁ 2 0 1 4 . ▁ ▁The ▁specific ▁ep ith et ▁t ac ana e ▁derived ▁from ▁the ▁T ac ana ▁vol cano , ▁which ▁is ▁on ▁the ▁border ▁of ▁Mexico ▁and ▁Gu atem ala , ▁and ▁the ▁second ▁highest ▁vol cano ▁in ▁Central ▁America . ▁ ▁Le pt oph ars a ▁t ac ana ▁is ▁the ▁first ▁la ce ▁bug ▁to ▁be ▁described ▁from ▁Mexican ▁am ber ▁foss ils , ▁while ▁the ▁related ▁Domin ican ▁am ber ▁fa una ▁is ▁much ▁more ▁diverse ▁with ▁six ▁described ▁species ▁as ▁of ▁ 2 0 1 4 : ▁E oc ader ▁bal yr uss us , ▁Le pt oph ars a ▁ev s y un ini , ▁Le pt oph ars a ▁fr ater , ▁Le pt oph ars a ▁po in ari , ▁Ste phan itis ▁roz