, ▁whereas ▁Le pt ol al ax ▁includes ▁all ▁the ▁species ▁from ▁Bor neo ▁( Le pt ol al ax ▁a ray ai , ▁Le pt ol al ax ▁dr ing i , ▁Le pt ol al ax ▁f rit in ni ens , ▁Le pt ol al ax ▁g rac ilis , ▁Le pt ol al ax ▁ham idi , ▁Le pt ol al ax ▁ma urus , ▁Le pt ol al ax ▁sab ah mont anus , ▁Le pt ol al ax ▁mar mor atus ▁and ▁Le pt ol al ax ▁p ict us ) ▁as ▁well ▁as ▁few ▁contin ental ▁species ▁( Le pt ol al ax ▁cro ce us , ▁Le pt ol al ax ▁mel icus ) ▁and ▁Le pt ol al ax ▁k aj ang ensis ▁from ▁Ti oman ▁Island , ▁at ▁least ▁provision ally . ▁ ▁Most ▁Le pt ol al ax ▁species ▁have ▁been ▁reported ▁from ▁very ▁small ▁areas ▁and ▁only ▁a ▁few ▁species ▁have ▁large ▁areas ▁of ▁occurrence . ▁Whether ▁this ▁distinction ▁is ▁real ▁or ▁the ▁w ides p read ▁species ▁represent ▁crypt ic ▁species ▁complex es ▁remains ▁to ▁be ▁confirmed . ▁ ▁Species ▁In ▁ 2 0 1 6 , ▁ 5 0 ▁species ▁were ▁known ▁in ▁the ▁genus . ▁New ▁species ▁are ▁still ▁being ▁described , ▁including ▁three ▁in ▁ 2 0 1 3 , ▁four ▁in ▁ 2 0 1 4 , ▁one ▁in ▁ 2 0 1 5 , ▁and ▁six ▁in ▁ 2 0 1 6 : ▁ ▁References