ides ▁collected ▁in ▁Maryland , ▁and ▁produced ▁fert ile , ▁all ot et rap lo id ▁off spring , ▁the ▁first ▁allo pol yp lo id ▁f ern s ▁to ▁be ▁artificial ly ▁produced ▁in ▁culture . ▁There ▁were ▁significant ▁morph ological ▁differences ▁between ▁these ▁artificial ly ▁produced ▁f ern s ▁and ▁those ▁from ▁Hav ana ▁Glen . ▁Wagner ▁and ▁Whit m ire ▁attributed ▁this ▁to ▁the ▁fact ▁that ▁the ▁pres umed ▁ancest ral ▁dip lo id ▁at ▁Hav ana ▁Glen ▁and ▁those ▁in ▁Maryland ▁had ▁origin ated ▁in ▁separate ▁hy brid ization ▁events ▁between ▁A .   pl aty neur on ▁and ▁A .   rh iz oph yll um ▁and ▁that ▁al lop lo id y ▁might ▁magn ify ▁gen etic ▁differences ▁between ▁the ▁parent al ▁species . ▁In ▁review ing ▁the ▁characteristics ▁of ▁the ▁species ▁in ▁ 1 9 8 2 , ▁Wagner ▁and ▁his ▁collabor ators ▁noted ▁that ▁as ▁a ▁hy brid , ▁the ▁name ▁of ▁the ▁species ▁was ▁more ▁correctly ▁written ▁As pl enium   ×   eben o ides , ▁according ▁to ▁the ▁International ▁Code ▁of ▁Botan ical ▁N omen cl ature . ▁However , ▁they ▁preferred ▁to ▁use ▁Wh er ry ' s ▁combination ▁of ▁× As pl en os orus ▁eben o ides . ▁( A ▁change ▁in ▁the ▁IC B N ▁in ▁ 1 9 7 2 ▁rendered ▁a ▁Latin ▁diagn osis ▁for ▁× As pl en os orus ▁unnecessary .) ▁Since ▁then , ▁ph y log en etic ▁studies ▁have ▁shown ▁that ▁Cam