family ▁was ▁originally ▁placed ▁in ▁family ▁C yst isc idae ▁by ▁Co over t ▁& ▁Co over t ▁( 1 9 9 5 ) ▁but ▁placed ▁back ▁in ▁Mar gin ell idae , ▁following ▁La ▁P erna ▁( 1 9 9 9 ) ▁and ▁based ▁on ▁the ▁morph ology ▁of ▁living ▁animals . ▁▁ ▁Gran ul ina ▁J ous se a ume , ▁ 1 8 8 8 : 1 9 1 ▁ ▁P ug n us ▁Hed ley , ▁ 1 8 9 6 : 1 0 5 - 1 0 6 ▁ ▁Pa ola ura ▁Sm ir ig lio ▁& ▁Mari ott ini , ▁ 2 0 0 1 ▁ ▁Sub family ▁MAR G INE LL ON IN AE ▁Co an ▁ 1 9 6 5 : 1 8 6 , 1 9 1 ▁ ▁Gen us ▁Af riv ol uta ▁Tom lin , ▁ 1 9 4 7 : 2 4 4 ▁Af riv ol uta ▁pr ing lei ▁Tom lin , ▁ 1 9 4 7 ; ▁O D ▁( M ) ▁Gen us ▁Mar gin ell ona ▁Mart ens , ▁ 1 9 0 4 ▁Mar gin ella ▁( M .) ▁gig as ▁Mart ens , ▁ 1 9 0 4 ; ▁( M ) ▁ ▁Sub family ▁MAR G INE LL IN AE ▁F lem ing , ▁ 1 8 2 8 : 3 2 8 ▁Gen us ▁C ari begin ella ▁Esp in osa ▁& ▁Or te a , ▁ 1 9 9 8 ▁Gen us