▁the ▁sub family ▁O xy c ory n inae . ▁But ▁while ▁they ▁are ▁highly ▁diverse , ▁they ▁are ▁not ▁particularly ▁spe ci ose , ▁with ▁nearly ▁ 2 0 ▁described ▁and ▁a ▁few ▁und es cribed ▁species ▁compared ▁to ▁the ▁more ▁than ▁ 1 5 0 ▁Hawai ian ▁species ▁of ▁Pro ter hin us ▁of ▁the ▁Ag ly cy der ini . ▁ ▁The ▁foss il ▁record ▁shows ▁that ▁the ▁three ▁O xy c ory n inae ▁tribes ▁were ▁well ▁distinct ▁by ▁the ▁mid - P ale ogene , ▁about ▁ 5 0 ▁million ▁years ▁ago ▁( my a ). ▁The ▁bel ids ▁as ▁a ▁whole ▁are ▁of ▁Jur ass ic ▁origin , ▁and ▁the ▁O xy c ory n ini ▁must ▁thus ▁have ▁evol ved ▁in ▁the ▁Late ▁C ret ace ous ▁or ▁perhaps ▁Pale oc ene , ▁roughly ▁some ▁ 1 0 0 - 6 0 ▁my a . ▁ ▁Description ▁The ▁adult ▁O xy c ory n ini ▁have ▁a ▁number ▁of ▁fairly ▁characteristic ▁tra its , ▁but ▁they ▁are ▁not ▁present ▁in ▁all ▁members ▁of ▁this ▁tribe . ▁Many ▁have ▁a ▁pron ot um ▁whose ▁h ind ▁margin ▁is ▁at ▁least ▁as ▁wide ▁as ▁the ▁el y tra ▁bases , ▁often ▁wider . ▁Ex cept ▁in ▁the ▁rather ▁primitive ▁O xy c ras ped us , ▁the ▁first ▁t ars om ere ▁ ▁is ▁smaller ▁than ▁the ▁second , ▁and ▁micro scop ically ▁the ▁lack ▁of ▁s per mat he ca , ▁with ▁the ▁s per mat he