c osa ▁group . ▁ ▁Black burn , ▁et ▁al . ▁( 2 0 1 3 ) ▁consider ▁Kal ou la ▁species ▁en demic ▁to ▁the ▁Philippines ▁to ▁form ▁a ▁mon oph y let ic ▁group , ▁containing ▁the ▁species ▁Kal ou la ▁w alter i , ▁Kal ou la ▁rig ida , ▁Kal ou la ▁conj unct a , ▁Kal ou la ▁p ict a , ▁Kal ou la ▁k aling ensis , ▁and ▁Kal ou la ▁k oka cii . ▁Kal ou la ▁likely ▁colon ized ▁the ▁Philippines ▁from ▁S out heast ▁Asia ▁during ▁the ▁Late ▁Mi oc ene , ▁and ▁then ▁radi ated ▁into ▁different ▁ec ot ypes . ▁K . ▁k aling ensis ▁and ▁K . ▁k oka cii ▁occup y ▁nich es ▁as ▁ar b ore al ▁tree - hole ▁fro gs ▁in ▁northern ▁and ▁southern ▁Lu zon ▁respectively , ▁while ▁K . ▁rig ida ▁and ▁K . ▁w alter i ▁are ▁ground ▁fro gs ▁in ▁northern ▁and ▁southern ▁Lu zon ▁respectively . ▁K . ▁p ict a , ▁which ▁is ▁a ▁terrest rial ▁ground ▁f rog , ▁and ▁K . ▁conj unct a , ▁which ▁is ▁sc ans orial ▁( cl im bing ) ▁and ▁pre fers ▁shr ubs , ▁have ▁established ▁themselves ▁throughout ▁the ▁Philipp ine ▁arch ip el ago . ▁K . ▁p ict a ▁as ▁well ▁as ▁K . ▁pul ch ra ▁display ▁the ▁lowest ▁internal ▁gen etic ▁divers ity , ▁but ▁also ▁have ▁w ides p read ▁ge ographical ▁distributions ▁that ▁may ▁have