. ▁Al ong ▁with ▁Or c utt ia ▁and ▁Ne ost ap f ia , ▁Tu ctor ia ▁is ▁one ▁of ▁three ▁gener a ▁in ▁the ▁tribe ▁Or c utt ie ae , ▁previously ▁out lined ▁by ▁Re eder ▁in ▁ 1 9 6 5 . ▁All ▁three ▁Tu ctor ia ▁species ▁were ▁formerly ▁assigned ▁to ▁Or c utt ia , ▁the ▁type ▁genus ▁of ▁Or c utt ie ae . ▁Re eder ▁ere cted ▁Tu ctor ia ▁after ▁determ ining ▁that ▁the ▁three ▁species ▁were ▁more ▁closely ▁related ▁among ▁themselves ▁than ▁to ▁any ▁of ▁the ▁other ▁Or c utt ia ▁species . ▁Shared ▁features ▁include ▁chrom os ome ▁numbers , ▁the ▁spir al ▁arrangement ▁of ▁sp ike lets ▁on ▁the ▁ra ch is , ▁the ▁absence ▁of ▁ju ven ile ▁leaves , ▁the ▁presence ▁of ▁lod ic ules , ▁and ▁the ▁presence ▁of ▁an ▁ep ib last ▁in ▁the ▁seed ▁emb ry os . ▁The ▁name ▁Tu ctor ia ▁is ▁an ▁an agram ▁of ▁Or c utt ia . ▁ ▁Rec ent ▁( 2 0 1 0 ) ▁mole cular ▁ph y log en etic ▁analysis ▁suggests ▁that ▁Tu ctor ia ▁is ▁not ▁mon oph y let ic ▁as ▁currently ▁circ ums cribed , ▁and ▁is ▁in ▁need ▁of ▁tax onom ic ▁revision . ▁ ▁Description ▁Tu ctor ia ▁species ▁have ▁their ▁sp ike lets ▁spir ally ▁arranged ▁on ▁the ▁axis ; ▁le mm as ▁are ▁entire ▁( with ▁a ▁smooth , ▁even ▁margin ) ▁or ▁d entic ulate ▁( fin ely