1 9 5 6 , ▁was ▁Ruth ▁S anger , ▁who ▁succeeded ▁him ▁in ▁the ▁post . ▁ ▁Career ▁In ▁ 1 9 3 7 , ▁after ▁training ▁at ▁St ▁Bar th ol ome w ’ s ▁Hospital ▁in ▁London , ▁Race ▁worked ▁as ▁a ▁ser ologist ▁in ▁the ▁blood - ty ping ▁department ▁being ▁established ▁by ▁Ron ald ▁Fish er ▁at ▁the ▁Gal ton ▁Labor atory ▁at ▁University ▁College ▁London . ▁The ▁ser um ▁unit ▁moved ▁to ▁Cambridge ▁at ▁the ▁beginning ▁of ▁the ▁Second ▁World ▁War , ▁and ▁in ▁ 1 9 4 1 ▁Race ▁and ▁Arthur ▁M our ant ▁began ▁investig ating ▁the ▁family ▁of ▁Rh ▁ant ig ens . ▁This ▁work ▁followed ▁from ▁develop ments ▁made ▁by ▁Karl ▁Land ste iner ▁and ▁Alexander ▁S . ▁Wiener ▁in ▁the ▁USA . ▁ ▁In ▁ 1 9 4 6 ▁Race ▁was ▁appointed ▁head ▁of ▁the ▁Medical ▁Research ▁Council ▁Blood ▁Group ▁Research ▁Unit . ▁During ▁the ▁same ▁year , ▁Ruth ▁S anger ▁moved ▁to ▁London ▁to ▁complete ▁her ▁Ph D . ▁She ▁joined ▁Race ' s ▁group ▁as ▁assistant ▁to ▁Race . ▁S anger ▁and ▁Race ▁married ▁in ▁ 1 9 5 6 ▁following ▁the ▁death ▁of ▁Race ' s ▁first ▁wife . ▁ ▁T ogether ▁Race ▁and ▁S anger ▁published ▁Blood ▁Gr oups ▁in ▁Man ▁in ▁ 1 9 5 0 , ▁which ▁eventually ▁sp anned ▁six ▁ed itions . ▁Their ▁work ▁continued ▁in ▁the ▁ 1 9 6 0 s ▁with ▁the ▁discovery ▁of ▁the ▁X g ▁ant igen ▁system ▁and