▁had ▁bother ed ▁to ▁travel ▁to ▁the ▁Sen cken berg ▁Museum ▁or ▁the ▁Über see - M useum ▁in ▁Germany ▁to ▁directly ▁compare ▁the ▁type ▁spec im ens ▁with ▁fro gs ▁from ▁Ok ina wa . ▁ ▁By ▁the ▁mid - 2 0 th ▁century , ▁it ▁was ▁recognized ▁that ▁on ▁both ▁Ok ina wa ▁and ▁the ▁Ya ey ama ▁group ▁brown ▁fro gs ▁could ▁be ▁found , ▁and ▁R . ▁ok in av ana ▁was ▁applied ▁to ▁them , ▁without ▁comparing ▁actual ▁spec im ens . ▁By ▁the ▁ 1 9 7 0 s , ▁it ▁was ▁realized ▁that ▁the ▁Ok ina wan ▁fro gs ▁were ▁not ▁found ▁on ▁the ▁Ya ey ama ▁Islands , ▁but ▁only ▁a ▁super f icial ly ▁similar ▁but ▁st outer ▁f rog ▁with ▁a ▁distinct ive ▁sup ra bra ch ial ▁g land ▁and ▁a ▁fol d ▁of ▁skin ▁running ▁from ▁the ▁shoulders ▁to ▁the ▁h ips . ▁The ▁Ya ey ama ▁fro gs ▁were ▁at ▁first ▁identified ▁as ▁R . ▁ad en ople ura ▁and ▁eventually ▁described ▁as ▁a ▁new ▁species , ▁R ana ▁ps alt es , ▁in ▁ 1 9 8 5 . ▁ ▁In ▁ 1 9 9 9 , ▁upon ▁exam ination ▁of ▁the ▁lect ot ype ▁of ▁Bo ett ger ' s ▁R . ▁ok in av ana , ▁it ▁was ▁realized ▁that ▁this ▁animal ▁was ▁very ▁similar ▁to ▁R . ▁ps alt es , ▁differ ing ▁from ▁the ▁Ok ina wan ▁brown ▁fro gs ▁just ▁as ▁the ▁recently ▁described