op one , ▁and ▁Pro top one ▁are ▁also ▁placed ▁in ▁Pon er inae . ▁ ▁The ▁genus ▁Cas ale ia ▁is ▁placed ▁in ▁Amb ly op on inae , ▁while ▁the ▁last ▁genus ▁Pse ude ct atom ma ▁is ▁in ▁E ct at omm inae . ▁ ▁Description ▁ ▁The ▁M esse le p one ▁queen ▁was ▁described ▁as ▁having ▁an ▁approximate ▁body ▁length ▁of ▁ ▁with ▁a ▁head ▁that ▁is ▁distinct ly ▁small , ▁the ▁mes os oma ▁being ▁almost ▁two ▁times ▁greater ▁in ▁length . ▁ ▁The ▁sub tri angular ▁mand ibles ▁are ▁up ▁to ▁one ▁half ▁the ▁head ▁length ▁and ▁have ▁to oth ed ▁che wing ▁faces . ▁ ▁Like ▁Pro top one ▁species , ▁the ▁anten na e ▁so ckets ▁are ▁widely ▁placed ▁on ▁the ▁head , ▁in ▁contrast ▁to ▁those ▁seen ▁in ▁P ach y cond y la ▁which ▁are ▁close ▁together . ▁The ▁node - sh aped ▁pet io le ▁is ▁shorter ▁than ▁the ▁height , ▁with ▁a ▁hel ci um ▁that ▁projects ▁of ▁the ▁front ▁face ▁of ▁the ▁g aster al ▁segment . ▁Both ▁the ▁queen ▁and ▁male ▁have ▁a ▁distinct ▁pet io le , ▁rect angular ▁in ▁side ▁view , ▁similar ▁to ▁some ▁species ▁of ▁Le pt ogen ys . ▁ ▁Both ▁g enders ▁also ▁have ▁a ▁g aster ▁with ▁the ▁first ▁segment ▁shorter ▁than ▁the ▁second ▁segment . ▁ ▁The ▁queen ▁has ▁eyes ▁that ▁are ▁o val ▁in ▁shape ▁and ▁placed ▁slightly ▁forward ▁from ▁the ▁heads ▁mid point . ▁ ▁The ▁g ena