▁Sk ull ▁ 4 , ▁like ▁the ▁other ▁D man isi ▁spec im ens , ▁has ▁been ▁interpreted ▁as ▁a ▁regional ▁variant ▁of ▁H . ▁ere ct us ▁sens u ▁l ato . ▁The ▁sk ull ▁displays ▁a ▁number ▁of ▁morph ological ▁land marks ▁which ▁identify ▁it ▁as ▁H . ▁ere ct us ▁or ▁H . ▁erg aster , ▁namely ▁the ▁project ing ▁bar - like ▁brow ▁rid ges , ▁sag itt al ▁ke eling , ▁and ▁fl atten ed ▁par iet als . ▁However , ▁it ▁also ▁shares ▁a ▁number ▁of ▁tra its ▁unique ▁to ▁the ▁D man isi ▁pale ode me , ▁such ▁as ▁its ▁relatively ▁small ▁c ran ial ▁capacity ▁which ▁most ▁closely ▁approxim ates ▁values ▁for ▁H . ▁hab ilis , ▁its ▁degree ▁of ▁mid fa cial ▁projection ▁and ▁front al ▁con str iction , ▁its ▁small ▁third ▁mol ars , ▁and ▁del icate ▁tym pan ic ▁plate . ▁The ▁suite ▁of ▁primitive ▁tra its ▁it ▁shares ▁with ▁earlier ▁H omo ▁and ▁Austral op it he cus ▁species ▁may ▁indicated ▁that ▁the ▁species ▁represents ▁the ▁transition ▁between ▁H . ▁hab ilis ▁and ▁H . ▁ere ct us , ▁however ▁its ▁similar ities ▁with ▁Asian ▁H . ▁ere ct us ▁provide ▁an ▁alternative ▁explanation ▁that ▁it ▁better ▁represents ▁ge ographic ▁isol ation ▁of ▁an ▁already ▁established ▁H . ▁ere ct us ▁species . ▁ ▁Similar ities ▁with ▁H omo ▁ere ct us ▁sens u ▁l ato ▁▁ ▁Project ing , ▁angular ▁brow ▁rid ges ▁ ▁E lev ated ▁nas al