, ▁they ▁completely ▁lost ▁the ▁digits ▁of ▁the ▁front ▁foot . ▁Advanced ▁tit an osa urs ▁had ▁no ▁digits ▁or ▁digit ▁b ones , ▁and ▁walked ▁only ▁on ▁horses ho e - sh aped ▁" st umps " ▁made ▁up ▁of ▁the ▁column ar ▁met ac arp al ▁b ones . ▁ ▁Print ▁evidence ▁from ▁Portugal ▁shows ▁that , ▁in ▁at ▁least ▁some ▁sau rop ods ▁( pro bably ▁bra ch ios aur ids ), ▁the ▁bottom ▁and ▁sides ▁of ▁the ▁fore foot ▁column ▁was ▁likely ▁covered ▁in ▁small , ▁sp iny ▁scales , ▁which ▁left ▁score ▁marks ▁in ▁the ▁prints . ▁In ▁tit an osa urs , ▁the ▁ends ▁of ▁the ▁met ac arp al ▁b ones ▁that ▁contact ed ▁the ▁ground ▁were ▁unus ually ▁broad ▁and ▁squ ared - off , ▁and ▁some ▁spec im ens ▁preserve ▁the ▁remains ▁of ▁soft ▁t issue ▁covering ▁this ▁area , ▁suggesting ▁that ▁the ▁front ▁feet ▁were ▁rim med ▁with ▁some ▁kind ▁of ▁padding ▁in ▁these ▁species . ▁ ▁Matthew ▁Bonn an ▁has ▁shown ▁that ▁sau rop od ▁din osa ur ▁long ▁b ones ▁grew ▁is omet r ically : ▁that ▁is , ▁there ▁was ▁little ▁to ▁no ▁change ▁in ▁shape ▁as ▁ju ven ile ▁sau rop ods ▁became ▁gig antic ▁adult s . ▁Bonn an ▁suggested ▁that ▁this ▁odd ▁scaling ▁pattern ▁( most ▁verte br ates ▁show ▁significant ▁shape ▁changes ▁in ▁long ▁b ones ▁associated ▁with ▁increasing ▁weight ▁support ) ▁might ▁be ▁related ▁to ▁a ▁st ilt - walk er ▁principle ▁( s ugg