▁mathematical ▁tal ents ▁and ▁ability ▁in ▁oper ational ▁research ." ▁ ▁Research ▁and ▁career ▁After ▁sp ending ▁nine ▁months ▁in ▁the ▁US ▁receiving ▁training ▁in ▁gen et ics ▁and ▁animal ▁bre eding ▁with ▁S ew all ▁Wright ▁and ▁Jay ▁Laur ence ▁L ush , ▁Robert son ▁took ▁up ▁his ▁work ▁at ▁N AB G RO , ▁based ▁in ▁Edinburgh . ▁The ▁organisation ▁was ▁split ▁into ▁two ▁parts , ▁and ▁Robert son ▁became ▁a ▁member ▁of ▁the ▁Unit ▁of ▁Animal ▁Gen et ics , ▁where ▁he ▁spent ▁the ▁rest ▁of ▁his ▁career . ▁At ▁first ▁he ▁worked ▁on ▁impro ving ▁da iry ▁cattle ▁using ▁mathematical ▁and ▁statistical ▁methods ▁to ▁dev ise ▁the ▁most ▁effective ▁bre eding ▁program mes . ▁A way ▁from ▁the ▁every day ▁work ▁of ▁the ▁Unit , ▁Robert son ▁also ▁worked ▁on ▁evolution ary ▁bi ology , ▁studying ▁how ▁variation ▁is ▁maintained ▁in ▁populations , ▁and ▁particularly ▁the ▁roles ▁of ▁mut ation ▁and ▁of ▁stabil ising ▁selection . ▁Hill ▁writes ▁that ▁Robert son ▁made ▁" original ▁contributions ▁to ▁the ▁theory ▁of ▁gen etic ▁change ▁in ▁small ▁populations ▁and ▁introduced ▁a ▁theory ▁of ▁limits ▁to ▁artificial ▁selection ▁… ▁a ▁combination ▁of ▁mathematical ▁insight , ▁quant itative ▁gen etic ▁principles , ▁and ▁practical ▁context , ▁of ▁which ▁only ▁he ▁was ▁capable ." ▁ ▁Robert son ▁continued ▁to ▁work ▁on ▁da iry - related ▁research ▁and ▁wider ▁theoretical ▁studies . ▁He ▁did ▁much ▁to ▁introduce ▁w ides p read ▁use ▁of ▁artificial ▁ins em ination ▁in ▁da iry ▁cattle , ▁and ▁worked ▁for ▁many