▁to ▁California . ▁In ▁ 1 8 9 9 ▁he ▁received ▁a ▁Ph . D . ▁at ▁the ▁University ▁of ▁California ▁with ▁a ▁dis sert ation ▁on ▁R hab d onia , ▁where up on ▁he ▁married ▁his ▁first ▁wife , ▁Anna ▁Maria ▁Land st rom . ▁The ▁couple ▁would ▁have ▁two ▁daughters : ▁Anna ▁Maria ▁and ▁Ol ga . ▁ ▁At ▁the ▁University ▁of ▁California , ▁he ▁was ▁promoted ▁to ▁assistant ▁professor ▁from ▁ 1 9 0 1 ▁to ▁ 1 9 0 8 ▁then ▁associate ▁professor ▁of ▁bot any ▁ 1 9 0 8 – 1 9 0 9 . ▁In ▁ 1 9 0 9 , ▁he ▁moved ▁to ▁Harvard ▁University ▁as ▁an ▁assistant ▁professor ▁of ▁bot any , ▁taking ▁a ▁step ▁down ▁in ▁rank ▁but ▁being ▁closer ▁to ▁the ▁important ▁Marine ▁Bi ological ▁Labor atory ▁at ▁Wood s ▁H ole . ▁He ▁rose ▁to ▁the ▁rank ▁of ▁professor ▁at ▁Harvard ▁in ▁ 1 9 1 3 ▁at ▁the ▁age ▁of ▁ 4 2 . ▁ ▁Sp ending ▁every ▁summer ▁at ▁the ▁labor atory ▁in ▁Wood s ▁H ole , ▁he ▁was ▁named ▁a ▁trust ee ▁in ▁ 1 9 1 9 . ▁He ▁was ▁elected ▁to ▁the ▁National ▁Academy ▁of ▁Sciences ▁in ▁ 1 9 1 9 . ▁From ▁ 1 9 1 9 ▁until ▁ 1 9 6 4 , ▁he ▁was ▁the ▁co - editor ▁of ▁the ▁Journal ▁of ▁General ▁Phys i ology , ▁along ▁with ▁founder ▁Jacques ▁Lo eb . ▁His ▁work ▁appeared ▁with ▁the ▁first