evidence: In late September 1838 a young man, aged 29, a former medical student and amateur naturalist, who had spent several years in the South Pacific studying plant and animal life, but who remained puzzled as to why "favourable variants" of each species survived while "unfavourable variants" were destroyed, sat perusing a book, as he later recalled, "for amusement." 1 The work which provoked Charles Darwin was T. R. Malthus's Essay on Population (1798), which he later claimed first suggested to him the idea that "on the whole the best fitted live." This idea Darwin would popularize through the notion of the "struggle for existence," a phrase which he famously claimed to use as a "metaphor" but which meant simply "the doctrine of Malthus applied with manifold force to the whole animal and vegetable kingdoms." 2 That application resulted in the publication [End Page 223] of the Origin of Species in 1859, an occasion hailed as "the greatest event of Queen Victoria's reign," 3 even "by far the most important ... in the history of the modern West." 4 It is well known, too, that Darwin's appreciation of Malthus was not unique even among naturalists.