query: parent_taxon protoscaphirhynchus
supports: Acipenser is a genus of sturgeons. With 17 species, many of which are threatened, it is the largest genus in the order Acipenseriformes. Pseudoscaphirhynchus is a genus of fishes in the Acipenseridae family. The three species, all of which are threatened, are restricted to Central Asia. The Maastrichtian is, in the ICS geologic timescale, the latest age (uppermost stage) of the Late Cretaceous epoch or Upper Cretaceous series, the Cretaceous period or system, and of the Mesozoic era or erathem. It spanned the interval from . The Maastrichtian was preceded by the Campanian and succeeded by the Danian (part of the Paleogene and Paleocene). The Triassic is a geologic period and system which spans 50.9 million years from the end of the Permian Period million years ago (Mya), to the beginning of the Jurassic Period Mya. The Triassic is the first period of the Mesozoic Era. Both the start and end of the period are marked by major extinction events. The Triassic was named in 1834 by Friedrich von Alberti, after the three distinct rock layers ("tri" meaning "three") that are found throughout Germany and northwestern Europered beds, capped by marine limestone, followed by a series of terrestrial mud- and sandstonescalled the "Trias". Huso is a genus of large sturgeons from Eurasia. It contains two species, which both are critically endangered: The Cenomanian is, in the ICS' geological timescale the oldest or earliest age of the Late Cretaceous epoch or the lowest stage of the Upper Cretaceous series. An age is a unit of geochronology: it is a unit of time; the stage is a unit in the stratigraphic column deposited during the corresponding age. Both age and stage bear the same name. The Paleogene (or ; also spelled Palaeogene or Palæogene; informally Lower Tertiary) is a geologic period and system that spans 43 million years from the end of the Cretaceous Period million years ago (Mya) to the beginning of the Neogene Period Mya. It is the beginning of the Cenozoic Era of the present Phanerozoic Eon. The Paleogene is most notable for being the time during which mammals diversified from relatively small, simple forms into a large group of diverse animals in the wake of the CretaceousPaleogene extinction event that ended the preceding Cretaceous Period.  Sturgeon is the common name for the 27 species of fish belonging to the family Acipenseridae. Their evolution dates back to the Triassic some 245 to 208 million years ago. The family is grouped into four genera: "Acipenser", "Huso", "Scaphirhynchus" and "Pseudoscaphirhynchus". Four species may now be extinct. Two closely related species, "Polyodon spathula" (paddlefish) and "Psephurus gladius" (Chinese paddlefish, possibly extinct) are of the same order, Acipenseriformes, but are in the family Polyodontidae and are not considered to be "true" sturgeons. Both sturgeons and paddlefish have been referred to as "primitive fishes" because their morphological characteristics have remained relatively unchanged since the earliest fossil record. Sturgeons are native to subtropical, temperate and sub-Arctic rivers, lakes and coastlines of Eurasia and North America. 
  Sturgeons are long-lived, late-maturing fishes with distinctive characteristics, such as a heterocercal caudal fin similar to that of sharks, and an elongated spindle-like body that is smooth-skinned, scaleless and armored with 5 lateral rows of bony plates called scutes. Several species can grow quite large, typically ranging 712 feet (2-3 m) in length. The largest sturgeon on record was a Beluga female captured in the Volga estuary in 1827, weighing and long. Most sturgeons are anadromous bottom-feeders which migrate upstream to spawn but spend most of their lives feeding in river deltas and estuaries. Some species inhabit freshwater environments exclusively while others primarily inhabit marine environments near coastal areas, and are known to venture into open ocean. The Cretaceous is a geologic period and system that spans 79 million years from the end of the Jurassic Period million years ago (Mya) to the beginning of the Paleogene Period Mya. It is the last period of the Mesozoic Era. The Cretaceous Period is usually abbreviated K, for its German translation "Kreide" (chalk). The Turonian is, in the ICS' geologic timescale, the second age in the Late Cretaceous epoch, or a stage in the Upper Cretaceous series. It spans the time between 93.9 ± 0.8 Ma and 89.8 ± 1 Ma (million years ago). The Turonian is preceded by the Cenomanian stage and underlies the Coniacian stage. Acipenseriformes is an order of basal ray-finned fishes that includes the sturgeons and paddlefishes, as well as some extinct families. Protoscaphirhynchus squamosus is an extinct sturgeon that lived from the Turonian to the Maastrichtian . Scaphirhynchus is a genus of sturgeons native to the United States of America. All species in this genus are considered to be threatened. The pallid sturgeon is Endangered and the Alabama sturgeon is Critically Endangered.
candidates: acipenser acipenseridae acipenseriformes bear huso mya neogene pseudoscaphirhynchus scaphirhynchus
answer: acipenseridae