He received his his BA is from Stanford University in 1993 and his M.D. from Harvard University in 1998; he completed a residency in internal medicine at Brigham and Women's Hospital, and was then a postdoctoral fellow at Whitehead Institute/MIT Center for Genome Research.
He received his his BS from Stanford University in 1993 and his M.D. from Harvard University in 1998. He completed his internship and residency in internal medicine at Brigham and Women's Hospital in Boston, and then pursued postdoctoral training at the Whitehead Institute/MIT Center for Genome Research.