E. a. actia, adusta, albigula, alpestris, alpina, ammophila, aphrasta, arcticola, argalea, atlas, balcanica, bicornis, brandti, chrysolaema, diaphora, elwesi, enthymia, enertera, flava, giraudi, hoyti, insularis, khamensis, kumerloevei, lactea, lamprochroma, leucansiptila, leucolaema, longirostris, merrilli, nigrifrons, oaxacae, occidentalis, penicillata, peregrina, praticola, przewalskii, rubea, sierrae, strigata, teleschowi hag utahensis,