Þis ȝetalu beinnaþ dēor þe ilċȝecyndisc ȝebǣre hæfþ, beinnende ȝehǣmende, ƿōgung, mōdlufu, ȝeocbend, and āċennendhād sƿā Bruce Bagemihl cȳðde in his 1999 bēċ Biological Exuberance: Animal Homosexuality and Natural Diversity.