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Years (LLS), 14.8 years (Leiden 85-plus study), and 11.4 years (Danish 1905 cohort), 73.2% (n = 691) (LLS), 84.8% (n = 828) (Leiden 85-plus study), and 98.9% (n = 1561) (Danish 1905 cohort) of the individuals had died. The data collection of the Netherlands Twin Register is supported by NWO (904-61-090; 904-61-193; 480-04-004; 400-05-717; SPI 56-464-14192), Center for Medical Systems Biology (NWO Genomics); Centre for Neurogenomics and Cognitive Research (CNCR-VU); the EU (EU/QLRT-2001-01254); Genome scan for neuroticism (NIMH R01 MH059160); the Geestkracht program of ZonMW (10-000-1002), and institutes involved in NESDA (VU University Medical Center, Leiden University Medical Center, GGZ Buitenamstel-Geestgronden, Rivierduinen, University Medical Center Groningen, GGZ Lentis, GGZ Friesland, GGZ Drenthe). The genotyping of the samples was provided through the Genetic Association Information Network (GAIN). Mortality analyses were performed with Stata 11.1 (StataCorp LP) using a sex-adjusted, left-truncated Cox proportional hazards model to adjust for late entry into the data set according to age. 

Author contributions: Serum parameter = b0 + b1*Age + b2*Sex + b3*(Age*Sex) + b4*Group + b5*rs429358 (Model 2). Authors contributed as follows: JD, MB, BTH, RGJW, PES; Data analysis, drafting manuscript, study design Leiden Longevity Study. H-WU, QH, JJH-D; Statistical data analysis. DK, RB, HEDS, NL, EBA, WMP, DH; Data collection and data analysis Leiden Longevity Study. MK, LC, HT, AJC, FR, EJdG, DIB, AGU, KC, CMD; Data analysis and data collection GWAS controls and replication cohorts. MP; DNA isolations Leiden Longevity Study. FJO, DAG; Data collection Leiden Longevity Study GWAS cases.

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Bertram L, Muenter MB, Mullin K, Blacker D, Tanzi RE. (2007) Systematic meta-analyses of Alzheimer disease genetic association studies: the AlzGene database. Nat. Genet., 39(1): 17-23.

Boerwinkle E, Utermann G. (1988) Simultaneous effects of the apolipoprotein E polymorphism on apolipoprotein E, apolipoprotein B, and cholesterol metabolism. Am. J. Hum. Genet., 42(1): 104-112.

Boomsma DI, Willemsen G, Sullivan PF, Heutink P, Meijer P, Sondervan D, Kluft C, Smit G, Nolen WA, Zitman FG, Smit JH, Hoogendijk WJ, van Dyck R., de Geus EJ, Penninx BW. (2008) Genome-wide association of major depression: description of samples for the GAIN Major Depressive Disorder Study: NTR and NESDA biobank projects. Eur. J. Hum. Genet., 16(3): 335-342.

Boyden SE, Kunkel LM. (2010) High-density genome-wide linkage analysis of exceptional human longevity identifies multiple novel loci. PLoS One, 5(8): e12432.

Christensen K, Johnson TE, Vaupel JW. (2006) The quest for genetic determinants of human longevity: challenges and insights. Nat. Rev. Genet., 7(6): 436-448.