नियासिननि आंखालनि अनगायैबो, पेलाग्राया प्रतिन आंखालजोंबो आखु-होजायो, आगु-गोदान गम-मायाव मोननै गुबै एमिन एसिडफोरनि गुदि आंखाल, लाइसिन आरो त्रिपतफान।
Besides the lack of niacin, pellagra was also characterized by protein deficiency, a result of the inherent lack of two key amino acids in pre-modern maize, lysine and tryptophan.