
carcinosomatidae

carcinosomatidae (paeru "carcinosoma" rakam nundi vachindi, antey "peetha sareeram") anedi euripterids yokka kutumbam, idhi antarinchipoyina jala keella samuuham. vaaru suupar family carcinosomatoidialo sabhyulu, dheeniki "carcinosoma" anie kooda paeru pettaru. carcinosomatids silaajaalu uttara america, eurup mariyu asealo kanugonabaddayi, eee kutumbam bahusha prapanchavyaaptamgaa pampineeni saadhinchindi, mariyu chivari ordovician nundi prarambha devonian varaku vayassu paridhiloo undhi. avi daadaapu puurtigaa samudra vaataavaranaala nundi telisina atyanta samudra euripteridlalo okati.
carcinosomatids parimaanamlo gananeeyamgaa maarutuu untaayi, jaatula podavu konni centimeters nundi telisina atipedda arthropods varaku. atipedda carcinosomatid jaathi, "carcinosoma panktatam", kaneesam podavuku cherukundi mariyu parimaanamlo annintikante pedda euripterid, "jackylopterus" kuu poteegaa nilichindi. padanirmaanaparamgaa, carcinosomatids itara euripteridla nundi chaalaa bhinnamgaa undevi, ivi vaati saktivantamaina mariyu vennemukala mundu bhagalu, vistruta mariyu gundrani kendra sareeram mariyu oka telson (sareeram yokka postereormost vibhajana) loe mugise sannani mariyu gottapu thokaku prasiddhi chendaayi, idhi saadhaaranamgaa edho oka vidhamgaa vakramgaa untundi. eee anusaranalatho, carcinosomatids thellatho chaalaa pooli untaayi, mariyu euripterids yokka saadhaarana paeru 'samudra teellu' gaa maaradaaniki eee samuuham dohadapadi undavacchu. eee kutumbamlo naalugu, bahusha aidu, jaatulu unnaih "carcinosoma", "ecarcinosoma", "eusarcana", "rhinocarcinosoma" mariyu bahusha samasyaatmaka jaathi "holmipterus".
carcinosomatids vaari kramabaddheekarinchani aakaraanni batti balamaina mariyu churukaina eethagaallu ayye avakaasam ledhu. vaaru nectobenthic (diguvaku sameepamlo eetha kottadam), bahusha agra maamsaahaarulu (vaati parimaanaanni batti) ledhaa scavengerlu, aaharam kosam travvadam ledhaa bahusha buradalo padi, aakasmika daadi chese mamsaharuluga vechi undatam vantivi kaavacchu.
vivarana.
carcinosomatid euripterids chaalaa jaatulu chaalaa peddavi ayinappatikee, jaathi mariyu jaatulanu batti parimaanamlo gananeeyamgaa bhinnamgaa undevi. atipedda jaathi "carcinosoma panktatam", idhi annintikante pedda euripteridlalo okati, konni namuunaalu idhi parimaanamlo atipedda euripterid ayina "jackylopterus" kuu poteegaa, podavuku kooda cherukundani suuchistunnaayi. atichinna carcinosomatid jaathi 4 centimeters (1.5 angulaalu) podavutho "eusarcana obesus".
padanirmaanasaastraparamgaa, carcinosomatids euripterids madhya chaalaa bhinnamgaa undevi. avi pedda eetha teddulu, saktivantamaina mariyu vennemukala mundu chetulu, vistruta mariyu gundrani prebdomen (kendra sareeram) mariyu sannani, gottapu pottikadupu, ivi tarachugaa vakramgaa unde variable padanirmaana shaastram yokka telson (sareeram yokka postereormost vibhajana) loe mugustaayi. oka konamlo, carcinosomatids kanipinchetappudu teella lantivi, mariyu euripterids yokka saadhaarana paeru 'samudra teellu' gaa maaradaaniki dohadapadi undavacchu.
samuuhamloo padanirmaana saastramlo gananeeyamaina vaividhyam undhi. samuuhamlooni sabhyulandarilo carapace tribhujaakaara nundi upachaturasraakaara aakaaramlo untundi, khachitamaina aakaram maaravacchu. rhinocarcinosoma" loe, carapace mundu bhaagamlo vilakshanhamaina, paara aakaaramlo podugu undedi. anni jaatulalo puurvapu padaardhaalu vistrutamgaa undevi, conny vaati vedalpu kooda jaatula nundi jaatulaku bhinnamgaa untundi. saapekshamgaa cheppalante, vistrutamaina jaathi "eusarcana obesus", dheenilo nalgava vibhagam modati enimidi vibhagalu kalipi podavugaa unnanta vedalpugaa undhi. spinocity (enni vennemukalu) mariyu mundu chetivella parimaanam kooda jaathi nundi jaatiki maarutuu untaayi, udaaharanaku "eusarcana" yokka mundu chetulu "rhinocarcinosoma" kante marinta saktivantamgaa untaayi. telson jaathi madhya gananeeyamgaa maarutuu untundih "rhinocarcinosoma" loe idhi balamgaa mariyu chadunainadi, konchem paiki vangi untundi, "eusarcana" loe idhi sthoopaakaaramlo untundi mariyu padunaina, teella lanty thoka spikega roopondinchabadindi mariyu "carcinosoma" loe idhi chadunainadi, itara euripteridlalo kanipinchani vistarinchina mariyu vibhajana nirmaanamloo mugustundi.
parisoedhana charitra.
1859loe zaan vilium salter "petrigotus punctatus" paerutoe varninchina mottamodati carcinosoma panktatam varninchabadina mottamodati carcinosomatid jaathi. taruvaata carcinosomatidga varninchabadina mottamodati jaathi "eusarcus" (mariyu daani rakam jaathi "ii. scarpionis), augustu r dwara varninchabadindi. newyork raashtramloni pridoli-yugam gede waterlime nundi swaadheenam chesukunna silaajaala aadhaaramgaa 1875loe groat mariyu vilium henry pitt. eee jaathi yokka vivarana lopinchindi mariyu silaajaala roopurekhalu mariyu aakaarampai matrame aadharapadi unnatlu anipinchindi, idhi henry woodward "ii" nii suuchimchadaaniki daariteesindi. euripterus punctatus" ("" petrigotus punctatus "" "" euripterus "" jaatigaa tirigi vargeekarinchabadindi) aakaaramlo samanamga undaalane kaaranamtho scarpionis "" euripterus "" kuu ". groat mariyu pitt laku teliyakundaane, "eusarcus" kuu appatike 1833loe gonyleptidae kutumbaaniki chendina (prastutam jeevinchi unna) laniatorid pantakotavaari jaatigaa paeru pettaru, mariyu idhi oka vyagramaina paeru. 1930la varaku eee paeru gurtinchabadaledu. 19va sataabdam chivaralo kooda varninchabadina "eurisoma" anae jaathi, daani rakam jaatulato paatu paeru pettabadindi, "e. 1890loe edward waller clay pole raasina newlini, 1890loe clay pole taruvaata aadhunika beatles yokka oka jaatiki chendinadani kanugonappudu, athanu "eurisoma" paerunu "carcinosoma" paerutoe bharti chesaadu.
1912loe, zaan masson clark mariyu rudolph rudeman "eusarcus" mariyu anni sambandhita rakaala euripteridla madhya thedaalu entha goppaga unnayante "eusarcus" itara euripteridla nundi bhinnamgaa undani "puurtigaa spashtamgaa" undani prakatinchaaru. clark mariyu rudeman aneka kotta jaatulanu "eusarcus" gaa paerkonnaaru, veetilo kotta jaatulu taruvaata "rhinocarcinosoma" jaatiki chendina jaatulugaa pariganinchabaddaayi, mariyu "eusarcus" "carcinosoma" kuu samanamga undani kooda nirdhaarinchaaru. carcinosoma" kante mundu "eusarcus" anie paeru pettabadinanduna, praadhaanyata yokka vargeekarana chattalu "eusarcus" anedi taxon paeru anie nirdesinchaayi.
"eusarcus" chivaraku 1934loe leaf starmer chetha oka vyagramaina paerugaa gurtinchabadindi. starmer tadupari andubaatulo unna athi puraatanamaina vyatirekhamaina paryaayapadamaina "carcinosoma" kuu paeru marchadu. 1934loe, "carcinosoma", "mixopterus", "ekinognathus" mariyu "megalograptus" anae naalugu jaatulanu kaligi undataaniki, starter carcinosomatidae kutumbaanni kooda pravesapettaadu, praarambhamlo 'carcinosomidae' paerutoe, 1934loe. eric enn dwara kutumbam savarinchabadindi. starmer yokka 1955 grandham "akasheruka polyontologypy kjelleswig-weiring" loe, paeru saraina carcinosomatidaga maarchabadindi mariyu "carcinosoma" kakunda itara jaatulu vaari swantha kutumbaalaku badilee cheyabaddaayi ("mixopterus" mixopteridae mariyu "megalograptus" mariyu "echinognathus" megalograptidaeku). 1942loe, embric strand enimidi samvatsaraala kritam starmer eee vishayanni parishkarinchinappatiki, "eusarcus", "eusarcana" kuu maroka pratyaamnaaya paerunu pratipaadinchindi. "rhinocarcinosoma" nuu 1962loe nester ivanovic novojilov "carcinosoma" nundi vaeru chesaaru, daani carapace itara carcinosoma kante bhinnamgaa undatam aadhaaramgaa ".
1964loe carcinosomatidsnu savarinchinappudu, kenneth edward caster mariyu eric enn. kjeleswig-weiring "eusarcus" mariyu "carcinosoma" lanu vibhina jaatulugaa gurtinchindi, 1912 paryaayapadaalu tappugaa unnayani nirnayinchindi. eusarcus" ninda, caster mariyu kjelleswig-weiring, bahusha strand yokka "eusarcana" gurinchi teliyaka, gatamlo "eusarcus" anie soochinche jaatulaku "paracarcinosoma" anae pratyaamnaaya paerunu roopondinchaaru. 1964loe kooda, caster mariyu kjelleswig-weiring "eusarcus"/"paracarcinosoma" yokka ordovician namuunaalanu parigananaloki teesukuni kotta jaatiki "ecarcinosoma" anie paeru pettaru. aaa namoonaalalo chaalaa varaku sudophasilsga gurtinchabadinappatiki, "ecarcinosoma" rakam namuunaa praamaanikamaina silaajam mariyu kutumbam yokka mottamodati rikaardu. 20va sataabdam chivaralo viyatnaamlo "rhinocarcinosoma" silaajaala aavishkaranatoe carcinosomatids yokka telisina bhougolika paridhi gananeeyamgaa vistarinchindi, dheeniki jaathi "aarr" anie paeru pettaru. 2002loe dosonances.
paracarcinosoma" nuu taruvaati parisoedhakulu tarachugaa upayoginchinappatiki, mundu paeru pettabadina "eusarcana" nuu jasson eah gurtinchaaru. dunlup mariyu james lansdell 2012loe "eusarcus" yokka chellubaatu ayye pratyaamnaaya paerugaa, "paracarcinosoma" kuu ketaayinchina jaatulanu aaa jaatiki badilee chesi, "paracarcinosoma" nuu juuniyar paryaayapadamgaa paerkonnaaru. 2015loe lansdell mariyu sahacharulu chesina phylogenetic vislaeshanhaloo "holmipterus" anae samasyaatmaka euripterid jaatini besal carcinosomatidga tirigi pondaaru. asampoorna shilaja padaartham mariyu vividha kutumbaala nundi lakshanaala spashtamaina kalayika kaaranamgaa "holmipterus" yokka sthaanam, euripterid kutumba vrukshamlo khachitamgaa ledhu mariyu daani silaajaalu rendu ververu jaatulanu kooda suuchimchavachchu, porapaatuna kalisi samuuhaparachabadi untaayi.
vargeekarana.
carcinosomatids infra arder diploperculata lopala suupar family carcinosomatoidialo bhaagamgaa vargeekarinchabaddaayi. carcinosomatoidialo mixopteridae mariyu megalograptidae kutumbaalu kooda unnaayi. carcinosomatidae gatamlo, 1989 nundi 2000 la prarambham varaku, suupar family 'hagmilarioidia' loe hagmileridae kutumbamtho samuuham cheyabadindi, vennemuka avayavalu mariyu anni avayavalu kaaranamgaa, eetha teddulu minaha, sthiramaina rakam. hagmileridae nedu suupar family petarigotioidiaku besal sabhyuluga pariganinchabadutondi.
carcinosomatoidia yokka antargata phylogeny sarigaa parishkarinchabadaledu (aspashtamgaa). krinda unna modati claudogram euripterid parisoedhakudu 2007 adhyayanam taruvaata undhi. eric tetley, idhi 2004 mariyu 2007 madhya nirvahinchina euripteridlapy vividha phylogenetic vislaeshanhala phalitaala aadhaaramgaa roopondinchabadindi, ayithe rendava claudogram james lansdell mariyu sahacharulu 2015loe chesina adhyayananni anusaristundi. rendu claudograms carcinosomatoidianu matrame pradarsinchadaaniki saralikrutam cheyabaddaayi. tetley (2007) carcinosomatidaenu paraphylletic groupinga tirigi pondindi, idhi carcinosomatoidia yokka besal sabhyulaku kaaranamavutundi, ayithe lansdell "mariyu itarulu".(2015) recovered the carcinosomatids as a monophyletic group.
tetley (2007)
lansdell "mariyu itarulu."(2015)
polyoacology.
carcinosomatid euripterids atyanta samudra euripteridslo okati, ivi okappudu dibbalu, konni madugu amarikalu mariyu lothaina neetilo unde nikshepaala nundi telisinavi. idhi lothaina jalaala nundi teliyani vaati sameepa bandhuvulaina mixopteridlaku viruddhamgaa untundi. lothaina jalaala nundi telisina ekaika itara euripterid kutumbam petrigotids, ivi carcinosomatids maadirigaane pampinhee cheyabaddaayi, ayithe ivi marinta vijayavantamayyaayi. terrigotids pampinhee aadhaaramgaa, carcinosomatids prapanchavyaaptamgaa unde avakaasam undhi. avi, petrigotidlatho paatu, silurian mariyu devonianlalo dakshina khandam gondwana nundi telisina ekaika euripterid kutumbam. samudram kaani nikshepaala nundi telisina ekaika carcinosomatid jaathi "rhinocarcinosoma" (dheenini samudra niksheepaalu anie kooda pilustaaru), idhi fluevial (nadhi) mariyu lacustrine (sarassu) amarikalalo kooda kanugonabadindi.
aneka itara eetha euripteridla sareeraala vale vaati sareeraalu kramabaddheekarinchabadananduna, mariyu vaati telsonla yokka pratyaekamaina padanirmaanaala kaaranamgaa, carcinosomatidlu chaalaa churukaina eethagaallu kakapovachhu, bahusha marinta nectobenthic (diguvaku sameepamlo eetha kottadam) jeevanasailini avalambinchavacchu. eee jeevanasaili mukhyamgaa "rhinocarcinosoma" loe udaaharanagaa cheppavacchu, ikkada daani carapace mundu bhaagamlo paara lanty podupu travvakam ledhaa "matti-rubbadam" kosam upayoginchabadi undavacchu mariyu itara carcinosomatidlatho poliste eetha teddulu parimaanamlo tagginchabaddaayi. vaati parimaanaanni batti, carcinosomatids agra maamsaahaarulu ledhaa scavengerlu kaavacchu, aaharam kosam thravvachhu ledhaa bahusha buradalu vesi, aakasmika daadi chese mamsaharuluga vechi undavacchu. vaaru purugulu, itara keellanoppulu, brachiopods mariyu chepalanu tini, vaari mundu avayavalanu upayoginchi aaharaanni notiloki nettavacchu.

