
tylopterella

vamshanasham sambhavicha jala aarthropodukalude oru koottamaaya yooripteridinte oru janussaanu tylopterella. otappettathum tharatthilumulla inathinte oru phoosil maathram, tii. kaanadayile elorayile avasana silurian kaalaghattatthile (ludlo yugam) nikshepangalil ninnu boyil "kandethi. nottu" ennartham varunna purathana greak padangalaaya τύλη (τερον), "chirak" ennartham varunna πτερον (πτερον) enniva chernnaanu eee janussinte peru rachichirikkunnathu. tylopterellayude maathruka kandethiya david boyaline bahumaanikkunna peril "boyil" enna inam ariyappedunnu.
carapase (prosomayude dorsal plate, thala) oru marginal riim kondu athirthi pankidunna, annukal vashathayi sthaapicha, aarr bhaagangalulla oru preabdoman, postabdoman (adivayarinte rund bhagangal), oru short spike polulla telson (ithu shareerathinte postariyormost divishanaayirunnu) ennivayulla ariyappedaatha oru janussaanith. motham neelam 7.5 centimeter (2.9 inch) aayi. eee savisheshathakal "tylopterella" yeei "onikkopterella", "alcanopterus" ennivaykkoppam onicopterellidae kudumbatthil sthaapikkunnu.
"tylopterella" athinte kattiyulla alankaarathinum shareerathinte uparithalathinum sradheyamaanu. athinte jodiyakkiya tuberkilukal allengil athinte randaamathethum anchaamathethumaaya bhaagangalude mukalilulla nobukal mattu pala yooripteraidukalil ninnum ithine vyathyasthamaakkunnu. athinte shareerathinte eee kanam uyarnna lavanaavasthakal moolamaanu, "tylopterella" nuu guelf roopeekaranavumaayi poruthappedendivannu; shakthippeduthiya shellukalulla mattu jeevikalum ithe sthalathu kandethiyittundu.
vivaranam.
mattu onikkopterellidukale, "tylopterella" oru cheria euripterid aayirunnu. ariyappedunna oreyoru maathrkayude motham valippam 7.5 centimeter (2.9 inch) maathramaanennu kanakkaakkappedunnu.
"tylopterella" ennathu ariyappedaatha oru janussaanu; "tii" yude tharam maathruka. shareerathinte pinbhagam samrakshikkunna "boyil", ithuvare athinte eaka recordine prathinidheekarikkunnu. kaarapesinu (prosomayude dorsal plate, thala) oru semioovettu (eathaandu oval) aakruthiyundaayirunnu, munvashath vrithaakruthiyilullathum (murichukondu churukkiyathum) pinbhaagathu churukkiyathum kaarapesinte vashangalil kooduthal uyarnnathum idungiyathumaaya oru rimminal athirthi pankidunnathumaayirunnu. neelathekkaal veethiyulla ithu uparithalathinte asaadhaaranamaaya kanathinum alankaarathinum shradheyamaayirunnu. 2 centimeter neelavum 2.7 centimeter veethiyumaayirunnu kaarapesinu. annukal erekkure vashathayi, renifom (beane aakruthiyilulla), ettavum valia vyaasamulla ekadesham 0.40 centimeter (0.16 inch) ullathum avaykkidayil 0.6 centimeter (0.13 inch) verthirikkappettathumaayinu. kannil ninnu thulyamaayi (athaayath, oru kannil ninnu matte kannil ninnu valare akale), oosellikal (lalithamaaya kannu polulla indriyangal) sthithicheyyunna oru cheria vrithaakrithiyilulla praadhaanyam undaayirunnu. kaarapesinte uparithalam graanulettu (granyoolukalulla) aayirunnu, koodaathe minitt vrithaakrithiyilulla kizhanguvargangal adangiya oru alankaaramundaayirunnu, chilath otappettathum mattullava rando moonno settukalaayi samyojippichathumaan. pradhana kaalsya nikshepangal (kaalsyam) ithu shakthippeduthi.
backie yooripteridukalilenne, opistosomaykku (adivire) panthrand bhagangalundaayirunnu. ithu preabdoman (segmentukal 1 muthal 6 vare), telescope cheytha (segmentukal parasparam ovarlaappu cheiyunna), postabdoman (segmentukal 7 muthal 12 vare) enningane vibhajichirikkunnu. ithu kattiyullathum cheruthum othukkamullathumaayirunnu, prathyekichu avasaanathe rund vibhagangalil. ithu oru calcario-chittinas (chittin) paali kondu moodappettirunnu. randaamathethu muthal anchaamathe vare terjaittukal (segmentinte dorsal pakuthi) oronnum athinte median lainil oru jodi valuthum kharavum pramukhavum neelamullathumaaya tuberkilukal allengil nobukal vahikkunnu. avaa adithattil punarroopakalppana cheyyappettathum mukalil oru paridhivare bailob (rund lobukal) ullathumaayirunnu. telson (shareerathinte ettavum pinbhaagathe vibhajanam) kramena idungiyathum cheruthaayi valanjathum chulivullathumaaya nattellu poleyaayirunnu. ithinu 1.5 centimeter (0.6 inch) neelamundaayirunnu. naalaamathe anubandhathinteyum (avayavathinteyum) neenthal kaalile paadilinteyum (aaraamatheyum avasaanatheyum jody anubandhangal) shakalangalum samrakshikkappettittund.
gaveshanathinte charithram.
"carapace, segmentukal, telson enniva samrakshikkunna orotta maathrkayaanu (holotype, gsc 2910, kaanadayile geological sarveyude myoosiyathil sookshichirikkunnu) tylopterella ennariyappedunnathu. britiish kammaran puraavastu gaveshakanumaaya david boyil, varshangalolam fossilukal shekharichukondu, 1881-ll kaanadayile elorayile guelf roopeekaranathilaanu ithu kandethiyathu. elora school myoosiyathile trasttikal (trastt sthaanathulla vyakthikal) kaanadayile geological sarveyude myoosiyathilekk phoosil konduvannu, avide britiish paaliyantolajistaaya joseph frederic vaittaves ithu vivarichu. yooripteras" enna janussinte oru puthiya inam addeham athinaayi nirmmichu, "iii. boyil ", athinte prathyeka peru athinte kandethalukaarane bahumaanikkunnu.
1912ll american paleantologistukalaaya joonn mesan clark rudolph rudeman chernnu "iii" ennu thiricharinju. tylopterus" enna swantham upavibhagam ullathinaal "yooripterasil" ninnu mathiyaya vyathyasthamaaya oru apakathakamaaya roopamaayi boyley ". pothuve hraswavum othukkamullathumaya body allengil "iii" yude kattiyulla calsues-chittinas integment polulla savisheshathakal ithinu anukoolamaayirunnu. boyil, "yooripterasil ninnu vyathyasthamayi, athil athu chittinas mathramaayirunnu. avar eee inathe mattu euripterid inangalaaya "wood wardopterous scabrosus", "hibertopterus steevansoni" ennivayumaayi thaarathamyam cheythu, iva randum "yooripterasinte" bhaagamaayirunnu, athil alankaaravum kattiyullathum kaalsyavumaayirunnu. anthraconectess" (innu "adelophthalmus" ennathinte paryaayamaayi amgeekarikkappedunnu) ennathinte gerontic (muthirnna) saambilukalilum eee swabhaavam kaanapedunnu, ennaal clark rudeman rund upavibhaagangalum thammilulla adutha bandham nishedhichu, kaaranam randum valare vyathyasthamaaya samudra saahacharyangalil jeevichirunnu ("aanthraakonektukal" uppuvellathilo shudhajalathilo jeevikkumbol "tylopterus" valare lavanajalathilo jeevichirunnu). tylopterus" enna peru purathana greak padangalaaya τύλη ("" tulay "," nottu "), πτερον (" "teron", "wing") ennivayaal rachikkappettittund, athinaal "" nottu wing "ennu vivarthanam cheyyappedunnu".
norwegian paleantologistum jiyolajistumaaya leaf stormar 1951-ll tylopterella ennathinu pakaram tylopterus enna peru maatti, kaaranam 1867-ll franch entomologist emm avatharippicha curculionid beettil janussaaya tylopterasinu ithu aadyam upayogicha oru peraayirunnu. gillom capiomont. "tylopterella" oru vyakthamaaya kaaranavumillathe "yooripterasil" ninnu oru prathyeka janussai shaasthreeya prabandhangalil prathyakshappedaan thudangum. 1958ll american paleantologist eric norman kelleswig-waring mattoru januss enna padavi aavarthichu urappikkunnathuvare tylopterella mattethoru euripterid janussil ninnum athinte savisheshamaaya alankaarathaal vyathyasthamaanennu avakashappettu.
1962ll stylonoorasinte oru inam. terjaittukalil jodiyakkiya kizhanguvargangal ullathine adisthaanamaakki russian paleantologist nesttar ivanovichu novojilov mennery tylopterellaykku niyogichu, ithu oru maathrkayil maathram samshayaaspadamaayi kaanapedunna oru swabhaavamaanu. menri ". ennirunnaalum, britiish paleantologist david jeey. yude neethrthwathilulla 2014 leeii padanatthilaanu eee inam punarnirmmichathu. marshell, ithu oru yooripteridine prathinidheekarikkunnille nirnnayichu, marichu "dwulikiaspis" ennu peritta oru puthiya chasmetaspidid janussine prathinidheekarikkunnu, angane "tylopterella" monotypic veendum avatharippikkunnu.
vargeekaranam.
"tylopterella" neey oonichooptterellide kudumbatthinte bhaagamaayi tharamthirickunnu, ithu monotypic suuppar famili oonichoopterellooyite eaka cled (group) aanu. athil orotta inam ulppedunnu, tii. boyil, "kaanadayile elorayude guelf roopeekaranathinte silurian ennathil ninnu.
yooripteras" enna inamaayi aadyam visheshippikkappetta tylopterella 1912-ll mattoru upavibhagamaayi amgeekarikkappettu. 1951-ll britiish professor scott simpson "drepanopteres abonances" enna tuberkilukale "tylopterella" enna tuberclukalumaayi thaarathamyam cheythu, ariyappedunna "tylopterella" enna eaka maathrkaye "drepanopteres", "stylonurus" ennivaye paraamarsikkaan kazhiyumennathinte adisthaanathil eee janussinte saadhuta vimukhathayode thiricharinju. kjelleswig-waring "tylopterella" yeei athintethaaya oru janussai thiricharinju, pakshe "eriopteras microphthalmus", "iii" ennivayumaayulla samanathakal kaaranam. eriensis, "addeham nirddeshichu" tii. boyley "eriopterasil" ninnu nerittu uthbhavichathaakam. "1989ll american paleantologist victor pii. aayirikkunnathuvare euripteridae kudumbatthinte bhaagamaayi tylopterellaye tharamthirichirunnu. apoorvamaaya vasthukkalum eathenkilum kudumbatthinte vyakthamaaya savisheshathakalude abhavavum kaaranam tollarton ithine "inserta sedis" (avyakthamaaya bandhangalulla oru taxon) ennu tharamthirichu.
ennirunnaalum, 2011-ll britiish jiyolajistum paleobiologistumaaya jaims sii. lamsdell puthiya suuppar famili onikkopterelloyidum onicopterellidae kudumbavum srishtichu, randaamathethinu "tylopterella", "onikkopterella" enniva nalki. randaamathethum naalaamathethumaaya jody anubandhangalil nattellukalude saannidhyam, anchaamatheyum aaraamatheyum nattellukalude abhavam (ettaamathe podomiyarile spurse ozhike, athaayath aaraamathe anubandhathinte legg segment ozhike), laattaral annukalulla kaarapesinte aakruthi, lansolate allengil styliphom telson ennivayaanu eee kudumbatthinte savisheshathakal. drepanopterus" "tylopterellayum" thammilulla eathenkilum sakhyam lamsdell nirasichu. "munbu karuthiyirunnathupole lalithamaaya oru projection (neendunilkkunna sareerabhaagam) ennathinekkaal neenthal kaalil chalikkunna nattellu kandethiyathinal moonnu varshathinu sesham oonikkoopterellideyk "alcanopterus" nalki.
thaazhe koduthirikkunna cladogram, mattu grouppukalumaayi yooripteridukalude yooripterina sub oordarile basel amgangale pradarshippikkunna lamsdell nadatthiya 2011 leeii phylogenetic vishakalanathile oru valia padanatthe (yooripteridukal maathram kaanikkan lalithamaakkunnath) adisthaanamaakkiyullathaanu.
palio ekkolaji.
kaanadayile ontaariyoyile ludlovian nikshepamaaya elorayil ninnaan "tylopterella" yude eaka phoosil kandeduthathu. tylopterella" yude shareera uparithalathinte angeyattatthe kanam kaaranam (kadalinekkaal lavanathayude alavu kooduthalulla) guelf roopeekaranathile janthujaalangal thurannukaattappedukayum pavizhapputtukalile jeevajaalangalkk nirantharam thiramaalakal adikkukayum cheiyunna hypersaline avasthakal aanennu karuthappedunnu. kattiyulla shellukalulla braakkiyopodukalumaayum molaskukalumaayum ithu undaakunnathum sushirangalulla parukkan daanyamulla dolomite "tylopterella" enna fosilukalude matrix ithu soochippikkunnu. athinaal, hypersalinatic anuyojyamaaya oreyoru yooripteridine "tylopterella" prathinidheekarikkunnu.
"tii. phoosil saittil kaanapedunna eaka yooripteridaanu boyley. troconematoid "loxoplocus solutta" allengil anomphaloid "pycnomphalus solarioids" polulla gastropod inangalumaayi ithu bandhappettirikkunnu, athupole thanne mattu pala anischithamaaya inam bewalves, bryosovans allengil sephalopodukal ennivayumaayi ithu bandhappettirikkunnu. athu jeevichirunna paristhithi aazhamillathathum upa alaikalullathumaaya (athaayath sooryaprakaasham samudrathinte adithattil athunnu) kadalayi kanakkaakkappedunnu, athinte litholaji (paarakalude bhauthika savisheshathakal) pradhaanamaayum petroleum saannidyamulla dolomite ulkkollunnu.

