 The Sclerodermatineae was first legitimately used by Manfred Binder and Andreas <unk> in 2002 based on molecular analyses of nuclear ribosomal large <unk> ( <unk> ) rRNA sequences from 60 species of Boletales . This research was an extension of Binder 's 1999 graduate work , in which he argued for the need to recognize the molecular differences of the <unk> fungi . Sclerodermatineae is one of six lineages of the Boletales recognized as a suborder ; the others are the <unk> , <unk> , <unk> , <unk> , and <unk> . Of the nine genera assigned to the Sclerodermatineae , three are <unk> ( Boletinellus , Gyroporus , and Phlebopus ) , and six are gasteroid ( Astraeus , Calostoma , Diplocystis , Pisolithus , and Scleroderma ) . Since the suborder 's original description , there have been several phylogenetic studies investigating the Sclerodermatineae . Some studies have revealed the existence of numerous cryptic species and have contributed to taxonomic expansion of the group . The " core " Sclerodermatineae include the genera Astraeus , Calostoma , Scleroderma , Pisolithus , Diplocystis , Tremellogaster ( all gasteroid ) , and the <unk> genus Gyroporus ; Phlebopus and Boletinellus resolved as sister to this core group . 
