 Another species complex in <unk> includes A. <unk> and related species . Henderson ( 1995 ) recognised A. <unk> and A. <unk> , considering the latter to either be an <unk> form of A. <unk> or a hybrid between it and A. <unk> . <unk> and <unk> accepted A. <unk> , but <unk> considered it a dubious taxon . <unk> <unk> was recognised as a distinct species by Michael <unk> and <unk> ; <unk> and <unk> and <unk> concurred , but Henderson considered it part of A. <unk> . <unk> also described a fourth member of this group , A. <unk> , and it is accepted by <unk> and <unk> . 
