 P. exclamans are haplodiploid insects , as are other Polistes species including Polistes <unk> , Polistes <unk> , and Polistes annularis . This means they have haploid males that produce identical haploid sperm , and diploid females that produce haploid eggs through meiosis . This has consequences for genetic relatedness within colonies , since sisters receive identical sperm if they share a father , and the normal half identical contribution from the mother . This results in sex @-@ biased conflicts between the queen and the workers with respect to the sex ratio , with workers preferring a more female biased sex ratio , and queens preferring equal investment in queens and males . 
