 Krause listed five features that indicate that UA 8699 is a marsupial , not a placental or primitive therian . There is a well @-@ developed <unk> at the outer back margin ( <unk> ) , between the hypoconid ( one of the main cusps ) and the hypoconulid ( a smaller cuspule ) . The hypoconulid itself is located far lingually , relatively far from the hypoconid . The trigonid and talonid are about as broad , the trigonid is low @-@ crowned , and wear is mainly horizontal , resulting in broad , flat exposed wear facets . Averianov and colleagues noted that zhelestids , a placental group , share all those traits , though to varying degrees , and that the hypoconulid in similarly aged marsupials is even more lingually located , " twinned " to the entoconid ( the cusp on the back lingual corner of the tooth ) . They wrote that the specimen is on the whole more similar to zhelestids than to marsupials . 
