 Aside from some papers published discussing myrmecophilous beetles and insect pests in forests , Clark predominately researched and published articles about the taxonomy of Australian ants . All of his articles and monographs were published in Australia . Throughout his career , Clark described around 200 different species of ants , but half of these may be synonyms . He worked and researched with all of the main groups of Australian ants , but his research and revisionary efforts were extensive with members of the former subfamily Cerapachyinae ( now Dorylinae ) , especially those of Phyracaces , the Myrmeciinae , Rhytidoponera and members of the tribe Dolichoderini . 
