 In 1996 , Kevin Thiele and Pauline Ladiges published a new arrangement for the genus , after cladistic analyses yielded a cladogram significantly different from George 's arrangement . With respect to B. verticillata , their findings largely accorded with George 's arrangement : section Oncostylis was discarded as polyphyletic , but series Spicigerae was inferred to be monophyletic , and B. verticillata appeared in a succession of clades with the species previously identified as its closest relatives : first B. littoralis , then B. seminuda , then B. brownii , and finally B. occidentalis ( red swamp banksia ) : 
