 The phylogenetic relationships determined by molecular techniques are not consistent with a classification scheme based on similarities in spore morphology . Several bolete taxa have been proposed to accommodate species with reddish @-@ brown to vinaceous @-@ brown , finely wrinkled ( rugulose ) to perforated or punctate spores ; these include Boletus subgenus Tylopilus , Tylopilus subgenus Porphyrellus , Austroboletus , or Porphyrellus . Austroboletus tristis and A. longipes , two species from Southeast Asia , share several spore characteristics with Spongiforma thailandica . All three species have spores with rugulose surfaces , sometimes with minute punctures , and the spores turn purple in potassium hydroxide . Additionally , the pores from the three species share similar surface ornamentation visible with scanning electron microscopy . In contrast to Spongiforma , however , Austroboletus spores do not have an apical pore . Genetic analysis shows that Spongiforma is more closely allied with the smooth @-@ spored Porphyrellus than with the rugulose @-@ punctate @-@ spored Austroboletus . 
