Holocene follows Pleistocene
Find the relation between <e1>Holocene<\e1> and <e2>Pleistocene<\e2>.
The Pleistocene coyote (Canis latrans orcutti), also known as the Ice Age coyote, is an extinct subspecies of coyote that lived in western North America during the Late Pleistocene era. Most remains of the subspecies were found in southern California, though at least one was discovered in Idaho. It was part of a carnivore guild that included other canids like foxes, gray wolves, and dire wolves. Compared to their modern Holocene counterparts, Pleistocene coyotes were larger and more robust, weighing, likely in response to larger competitors and prey rather than Bergmann's rule. Their skulls and jaws were significantly thicker and deeper than in modern coyotes, with a shorter and broader rostrum and wider carnassial (denoting the large upper premolar and lower molar teeth of a carnivore, adapted for shearing flesh) teeth. These adaptions allowed it to cope with higher levels of stress, when it killed larger prey, compared to modern coyotes. Pleistocene coyotes were also likely more specialized carnivores than their descendants, as their teeth were more adapted to shearing meat, showing fewer grinding surfaces which were better suited for processing vegetation. The lower jaw was also deeper, and the molars showed more signs of wear and breakage than modern populations, thus indicating that the animals consumed more bone than today. Behaviorally, it is likely to have been more social than the modern coyote, as its remains are the third most common in the La Brea Tar Pits, after dire wolves and sabre - toothed cats, both thought to be gregarious species. Their reduction in size occurred within 1,000 years of the occurrence of the Quaternary extinction event, when the climate changed and the majority of their larger prey became extinct. Furthermore, Pleistocene coyotes were unable to successfully exploit the big game hunting niche left vacant after the extinction of the dire wolf, as that gap was rapidly filled by gray wolves. These gray wolves are likely to have actively killed off the larger - bodied coyotes, with natural selection favoring the modern gracile morph. Human predation on the Pleistocene coyote's dwindling prey base may have also impacted the animal's change in morphology. 
Holocene Pleistocene