title: Harpalus
text: # Harpalus
  
  Harpalus es un género de coleópteros adéfagos de la familia Carabidae.
  Principalmente holártico con unas pocas especies en la región afrotropical. Hay 415 especies.
  
  # Harpalus
  ## Algunas especies
  
  Comprende las siguientes especies:
  * Harpalus actiosus Casey, 1914
  * Harpalus acupalpoides Reitter, 1900
  * Harpalus adenticulatus Huang, 1992
  * Harpalus aeneipennis Faldermann, 1836
  * Harpalus aenigma (Tschitscherine, 1897)
  * Harpalus aequicollis Motschulsky, 1844
  * Harpalus aesculanus Pantel, 1888
  * Harpalus affinis (Schrank, 1781)
  * Harpalus agakhaniantzi Mikhailov, 1972
  * Harpalus akinini Tschitscherine, 1895
  * Harpalus alajensis Tschitscherine, 1898
  * Harpalus albanicus Reitter, 1900
  * Harpalus alexandrae Kataev, 1990
  * Harpalus alexeevi Kataev, 1990
  * Harpalus alienus Bates, 1878
  * Harpalus alpivagus Tschitscherine, 1899
  * Harpalus amarellus Bates, 1891
  * Harpalus amariformis Motschulsky, 1844
  * Harpalus amplicollis Menetries, 1848
  * Harpalus amputatus Say, 1830
  * Harpalus anatolicus Tschitscherine, 1898
  * Harpalus angulatus Putzeys, 1878
  * Harpalus angustipennis Boheman, 1848
  * Harpalus angustitarsis Reitter, 1887
  * Harpalus animosus Casey, 1924
  * Harpalus anisodactyliformis Solsky, 1874
  * Harpalus antonowi Tschitscherine, 1898
  * Harpalus anxioides Kataev, 1991
  * Harpalus anxius (Duftschmid, 1812)
  * Harpalus aogashimensis (Habu, 1957)
  * Harpalus apache Kataev, 2010
  * Harpalus araraticus Mlynar, 1979
  * Harpalus arcuatus Tschitscherine, 1898
  * Harpalus arnoldii Kataev, 1988
  * Harpalus ascetes Kataev & Wrase, 1997
  * Harpalus asemus Basilewsky, 1946
  * Harpalus asphaltinus Roth, 1851
  * Harpalus aterrimus Casey, 1914
  * Harpalus atratus Latreille, 1804
  * Harpalus atrichatus Hatch, 1949
  * Harpalus atripes Casey, 1914
  * Harpalus attenuatus Stephens, 1828
  * Harpalus australasiae Dejean, 1829
  * Harpalus autumnalis Duftschmid, 1812
  * Harpalus azumai Habu, 1968
  * Harpalus babai Habu, 1973
  * Harpalus badakschanus Jedlicka, 1956
  * Harpalus baleensis Clarke, 1973
  * Harpalus balli Noonan, 1991
  * Harpalus basanicus J.R. Sahlberg, 1911
  * Harpalus basilewskyi Facchini, 2003
  * Harpalus basuto Basilewsky, 1958
  * Harpalus bellieri Reiche, 1861
  * Harpalus beneshi Kataev & Wrase, 1997
  * Harpalus bitinctus Jeannel, 1948
  * Harpalus brachypterus Tschitcherine, 1898
  * Harpalus brevicornis Germar, 1824
  * Harpalus brevis Motschiulsky, 1844
  * Harpalus breviusculus Chaudoir, 1846
  * Harpalus brunnipes Dejean, 1829
  * Harpalus bucharicus Tschitscherine, 1898
  * Harpalus bungii Chaudoir, 1844
  * Harpalus caeruleatus Bates, 1878
  * Harpalus caiphus Reiche & Saulcy, 1855
  * Harpalus calathoides Motschulsky, 1844
  * Harpalus calceatus (Duftschmid, 1812)
  * Harpalus caliginosus (Fabricius, 1775)
  * Harpalus capicola Dejean, 1829
  * Harpalus capito A.Morawitz, 1862
  * Harpalus cardoni Antoine, 1922
  * Harpalus caspius (Steven, 1806)
  * Harpalus cautus Dejean, 1829
  * Harpalus cephalotes Fairmaire & Laboulbene, 1854
  * Harpalus chalcentus Bates, 1873
  * Harpalus chalcopterus Jeannel, 1948
  * Harpalus chasanensis Lafer, 1989
  * Harpalus chengjiangensis Huang, 1993
  * Harpalus chiloschizontus Huang, 1995
  * Harpalus chobautianus Lutshnik, 1922
  * Harpalus chrysopus Reitter, 1887
  * Harpalus cilihumerus Huang, Hu & Sun, 1994
  * Harpalus circumpunctatus Chaudoir, 1846
  * Harpalus cisteloides Motschulsky, 1844
  * Harpalus cohni Ball, 1972
  * Harpalus colchicus Motschulsky, 1850
  * Harpalus compar Leconte, 1848
  * Harpalus compressus Motschulsky, 1844
  * Harpalus contemptus Dejean, 1829
  * Harpalus convexus Faldermann, 1836
  * Harpalus cordifer Notman, 1919
  * Harpalus cordifer Notman, 1919
  * Harpalus coreanus (Tschitscherine, 1895)
  * Harpalus corporosus Motschulsky, 1861
  * Harpalus corpulentus (Casey, 1914)
  * Harpalus corrugatus Basilewsky, 1958
  * Harpalus crates Bates, 1883
  * Harpalus cupreus Dejean, 1829
  * Harpalus cursorius Peringuey, 1896
  * Harpalus cyanopterus Tschitscherine, 1897
  * Harpalus cyclogonus Chaudoir, 1844
  * Harpalus cyrtonotoides Notman, 1919
  * Harpalus danieli Reitter, 1900
  * Harpalus davidi (Tschitscherine, 1897)
  * Harpalus davidianus Tschitscherine, 1903
  * Harpalus decipiens Dejean, 1829
  * Harpalus defector Peringuey, 1896
  * Harpalus demelti Korge, 1962
  * Harpalus desertus Leconte, 1859
  * Harpalus diligens Tschitscherine, 1898
  * Harpalus dimidiatus (P. Rossi, 1790)
  * Harpalus disaogashimensis Huang In Huang & Zhang, 1995
  * Harpalus disimuciulus Huang, Lei, Yan & Hu, 1996
  * Harpalus dispar Dejean, 1829
  * Harpalus dissitus Antoine, 1931
  * Harpalus distinguendus (Duftschmid, 1812)
  * Harpalus diversicollis Basilewsky, 1958
  * Harpalus dubius Boheman, 1848
  * Harpalus durangoensis Bates, 1891
  * Harpalus ebeninus Heyden, 1870
  * Harpalus eberlovi O. Berlov, 1996
  * Harpalus egenus Dejean, 1829
  * Harpalus egorovi Lafer, 1989
  * Harpalus ellipsis Leconte, 1848
  * Harpalus eous Tschitscherine, 1901
  * Harpalus eraticus Say, 1823
  * Harpalus erosus Mannerheim, 1825
  * Harpalus erythropus Dejean, 1829
  * Harpalus euchlorus Menetries, 1839
  * Harpalus exiguus Boheman, 1848
  * Harpalus famelicus Tschitscherine, 1898
  * Harpalus farkaci Kataev & Wrase, 1995
  * Harpalus faunus Say, 1823
  * Harpalus ferghanensis Tschitscherine, 1899
  * Harpalus fimetarius Dejean, 1829
  * Harpalus flavescens Piller & Mitterpacher, 1783
  * Harpalus flavicornis Dejean, 1829
  * Harpalus fokienensis Schauberger, 1930
  * Harpalus foveiger Tschitscherine, 1895
  * Harpalus franzi Mateu, 1954
  * Harpalus frater Chaudoir, 1876
  * Harpalus fraternus Leconte, 1852
  * Harpalus froelichii Sturm, 1818
  * Harpalus fukiensis Jedlicka, 1956
  * Harpalus fulgens Csiki, 1932
  * Harpalus fulvicornis Thunberg, 1806
  * Harpalus fulvilabris Mannerheim, 1853
  * Harpalus fulvipennis Chaudoir, 1843
  * Harpalus fulvus Dejean, 1829
  * Harpalus fuscicornis Menetries, 1832
  * Harpalus fuscipalpis Sturm, 1818
  * Harpalus fuscipennis Wiedemann, 1825
  * Harpalus fuscoaeneus Dejean, 1829
  * Harpalus ganssuensis Semenov, 1889
  * Harpalus giacomazzoi Kataev & Wrase, 1996
  * Harpalus gilgil Basilewsky, 1946
  * Harpalus gisellae Csiki, 1932
  * Harpalus glasunovi Kataev, 1987
  * Harpalus gravis Leconte, 1858
  * Harpalus gregoryi Alluaud, 1917
  * Harpalus grilli Kataev, 2001
  * Harpalus griseus (Panzer, 1796)
  * Harpalus hartmanni Kataev, 2001
  * Harpalus hatchi Ball & Anderson, 1962
  * Harpalus hauserianus Schauberger, 1929
  * Harpalus herbivagus Say, 1823
  * Harpalus heyrovskyi Jedlicka, 1928
  * Harpalus hiekei Kataev & Wrase, 2010
  * Harpalus hirtipes Panzer, 1796
  * Harpalus honestus (Dufischmid, 1812)
  * Harpalus hospes Sturm, 1818
  * Harpalus huachuca Ball, 1972
  * Harpalus hybridus Boheman, 1848
  * Harpalus hypogeomysis Huang, 1993
  * Harpalus idiotus Bates, 1889
  * Harpalus imerinae Jeannel, 1948
  * Harpalus impressicollis Jeannel, 1948
  * Harpalus impressus Roth, 1851
  * Harpalus inconcinnus Chaudoir, 1876
  * Harpalus indianus Csiki, 1932
  * Harpalus indicola Bates, 1878
  * Harpalus indicus Bates, 1891
  * Harpalus indigens Casey, 1924
  * Harpalus inexspectatus Kataev, 1989
  * Harpalus ingenuus Tschitscherine, 1898
  * Harpalus innocuus Leconte, 1863
  * Harpalus italus Schaum, 1860
  * Harpalus janinae Jeanne, 1984
  * Harpalus jeanneli Basilewsky, 1946
  * Harpalus jordanus Jedlicka, 1964
  * Harpalus jureceki (Jedlicka, 1928)
  * Harpalus kabakianus Kataev, 1988
  * Harpalus kabakovi Kataev, 1987
  * Harpalus kadleci Kataev & Wrase, 1995
  * Harpalus kadyrbekovi Kataev, 1988
  * Harpalus kagyzmanicus Kataev, 1984
  * Harpalus kailiensis Huang, 1992
  * Harpalus kandaharensis Jedlicka, 1956
  * Harpalus karamani Apfelbeck, 1902
  * Harpalus karokorum Jedlicka, 1958
  * Harpalus katiae F.Battoni, 1985
  * Harpalus kazanensis Jedlicka, 1958
  * Harpalus kaznakovi Kataev & Wrase, 1997
  * Harpalus kibonoti Alluaud, 1926
  * Harpalus kirgisicus Motschulsky, 1844
  * Harpalus kiritshenkoi Kataev, 1990
  * Harpalus klapperichi Jedlicka, 1956
  * Harpalus kmecoi Facchini, 2003
  * Harpalus koshiensis Kirschenhofer, 1992
  * Harpalus kozlovi Kataev, 1993
  * Harpalus kryzhanovskii Kataev, 1988
  * Harpalus kunarensis Kataev, 1993
  * Harpalus laetus Reiche, 1843
  * Harpalus laevipes Zetterstedt, 1828
  * Harpalus lama Kataev & Wrase, 1997
  * Harpalus lampronotus Jeannel, 1948
  * Harpalus lateralis Dejean, 1829
  * Harpalus laticeps Leconte, 1850
  * Harpalus latus (Linnaeus, 1758)
  * Harpalus lederi Tschitscherne, 1899
  * Harpalus lethierryi Reiche, 1860
  * Harpalus lewisii Leconte, 1865
  * Harpalus liobasis Chaudoir, 1868
  * Harpalus litigiosus Dejean, 1829
  * Harpalus longicollis Leconte, 1848
  * Harpalus longihornus Lei & Huang, 1997
  * Harpalus longipalmatus Mordkovitsh, 1969
  * Harpalus lopezi Serrano & Lencina, 2008
  * Harpalus lugubris Boheman, 1848
  * Harpalus lumbaris Mannerheim, 1825
  * Harpalus luteicornis Dufischmid, 1812
  * Harpalus lutshniki Schauberger, 1932
  * Harpalus macronotus Tschitscherine, 1893
  * Harpalus madagascariensis Dejean, 1831
  * Harpalus mairei Peyerimhoff, 1928
  * Harpalus major Motschulsky, 1850
  * Harpalus makhekensis Basilewsky, 1958
  * Harpalus manas Kataev, 1990
  * Harpalus marginellus Gyllenhal, 1827
  * Harpalus mariae Kataev, 1997
  * Harpalus martini Vandyke, 1926
  * Harpalus masoreoides Bates, 1878
  * Harpalus massarti Burgeon, 1935
  * Harpalus mauritanicus Gaubil, 1844
  * Harpalus medvedevi Kataev, 2006
  * Harpalus megacephalus Leconte, 1848
  * Harpalus meghalayensis Kataev, 2001
  * Harpalus melancholicus Dejean, 1829
  * Harpalus melaneus Bates, 1878
  * Harpalus meridianus Andrewes, 1923
  * Harpalus metallinus Menetries, 1836
  * Harpalus metarsius Andrewes, 1930
  * Harpalus meteorus Basilewsky, 1946
  * Harpalus michaili Kataev, 1990
  * Harpalus michailovi Kataev, 1987
  * Harpalus microdemas Schauberger, 1932
  * Harpalus microthorax Motschulsky, 1849
  * Harpalus miles Peringuey, 1896
  * Harpalus minutulus Kataev & Liang, 2004
  * Harpalus mitridati Pliginskiy, 1915
  * Harpalus mlynari Kataev, 1990
  * Harpalus modestus Dejean, 1829
  * Harpalus morvani Kataev, 2001
  * Harpalus muciulus Huang, 1992
  * Harpalus nanniscus Peringuey, 1896
  * Harpalus natalensis Boheman, 1848
  * Harpalus natalicus Peringuey, 1896
  * Harpalus neglectus Audinet-Serville, 1821
  * Harpalus nevadensis K. Daniel & J. Daniel, 1898
  * Harpalus nigrans A. Morawitz, 1862
  * Harpalus nigritarsis C.R. Sahlberg, 1827
  * Harpalus numidicus Bedel, 1893
  * Harpalus nyassicus Basilewsky, 1946
  * Harpalus obliquus G.Horn, 1880
  * Harpalus oblitus Dejean, 1829
  * Harpalus obnixus Casey, 1924
  * Harpalus obtusiusculus Jeannel, 1948
  * Harpalus ochropus Kirby, 1837
  * Harpalus oodioides Dejean, 1829
  * Harpalus opacipennis (Haldemann, 1843)
  * Harpalus optabilis Dejean, 1829
  * Harpalus ovtshinnikovi Kataev, 1990
  * Harpalus pachys Jeannel, 1948
  * Harpalus paivanus Wollaston, 1867
  * Harpalus pallidipennis A. Morawitz, 1862
  * Harpalus pallipes Chaudoir, 1837
  * Harpalus parasinuatus Kataev & Liang, 2007
  * Harpalus paratus Casey, 1924
  * Harpalus parvulus Dejean, 1829
  * Harpalus pastor Motschulsky, 1844
  * Harpalus penglainus Huang, Hu & Sun, 1994
  * Harpalus pensylvanicus (Degeer, 1774)
  * Harpalus periglabellus Huang, 1992
  * Harpalus perlucens Bates, 1878
  * Harpalus petri Tschitscherine, 1902
  * Harpalus picipennis Duftschmid, 1812
  * Harpalus pilosus Huang, 1995
  * Harpalus plancyi Tschitscherine, 1897
  * Harpalus platynotus Bates, 1873
  * Harpalus plenalis Casey, 1914
  * Harpalus politus Dejean, 1829
  * Harpalus polyglyptus Schaum, 1862
  * Harpalus poncei Will, 2001 Usa (Florida)
  * Harpalus potanini Tschitscherine, 1906
  * Harpalus poussereaui Pierre & Mauro, 2010
  * Harpalus praecurrens Schauberger, 1934
  * Harpalus praticola Bates, 1891
  * Harpalus procognatus Lorenz, 1998
  * Harpalus progrediens Schauberger, 1922
  * Harpalus prosperus Basilewsky, 1972
  * Harpalus protractus Casey, 1914
  * Harpalus providens Casey, 1914
  * Harpalus pseudohauserianus Kataev, 2001
  * Harpalus pseudophonoides Schauberger, 1930
  * Harpalus pseudoserripes Reitter, 1900
  * Harpalus pseudotinctulus Schauberger, 1932
  * Harpalus pterostichus Reitter, 1900
  * Harpalus puetzi Kataev & Wrase, 1997
  * Harpalus pulchirinulus Reitter, 1900
  * Harpalus pulvinatus Menetries, 1848
  * Harpalus pumilus Sturm, 1818
  * Harpalus punctatostriatus Dejean, 1829
  * Harpalus puncticeps (Casey, 1914)
  * Harpalus punctipennis Mulsant, 1852
  * Harpalus pusillus Motschulsky, 1850
  * Harpalus pygmaeus Dejean, 1829
  * Harpalus quadratus Chaudoir, 1846
  * Harpalus raphaili Kataev, 1997
  * Harpalus reflexus Putzeys, 1878
  * Harpalus regressus (Casey, 1914)
  * Harpalus remboides Solsky, 1874
  * Harpalus retractus Leconte, 1863
  * Harpalus reversus Casey, 1924
  * Harpalus rivalsi Jeannel, 1948
  * Harpalus roninus Bates, 1873
  * Harpalus rotundus Facchini, 2003
  * Harpalus rougemonti Clarke, 1973
  * Harpalus rubefactus Bates, 1873
  * Harpalus rubripes Dufischmid, 1812
  * Harpalus rufipalpis Sturm, 1818
  * Harpalus rufipes (Degeer, 1774)
  * Harpalus rufiscapus Gebler, 1833
  * Harpalus rufitarsoides Schauberger, 1934
  * Harpalus rufomarginatus (Boheman, 1848)
  * Harpalus rumelicus Apfelbeck, 1904
  * Harpalus salinulus Reitter, 1900
  * Harpalus salinus Dejean, 1829
  * Harpalus sanctaehelenae Basilewsky, 1972
  * Harpalus saxicola Dejean, 1829
  * Harpalus scabripectus Huang, Hu & Sun, 1994
  * Harpalus schaumii Wollaston, 1864
  * Harpalus semenowi Tschitscherine, 1901
  * Harpalus sericatus Tschitscherine, 1906
  * Harpalus serripes Quensel, 1806
  * Harpalus servus Duftschmid, 1812
  * Harpalus setiporus Reitter, 1894
  * Harpalus seyrigi Jeannel, 1948
  * Harpalus siculus Dejean, 1829
  * Harpalus signaticornis (Duftschmid, 1812)
  * Harpalus silipes Dejean, 1831
  * Harpalus simplicidens Schauberger, 1929
  * Harpalus singularis Tschitscherine, 1906
  * Harpalus sinicus Hope, 1845
  * Harpalus sinuatipennis Jeannel, 1948
  * Harpalus sinuatus Tschitscherine, 1893
  * Harpalus smaragdinus Duftschmid, 1812
  * Harpalus smyrnensis Heyden, 1888
  * Harpalus solitaris Dejean, 1829
  * Harpalus somereni Basilewsky, 1946
  * Harpalus somnulentus Dejean, 1829
  * Harpalus spadiceus Dejean, 1829
  * Harpalus spretus Peringuey, 1896
  * Harpalus spurius Peringuey, 1896
  * Harpalus stephani Ball, 1972
  * Harpalus stevenii Dejean, 1829
  * Harpalus stoetznerianus Schauberger, 1932
  * Harpalus strenuus Tschitscherine, 1898
  * Harpalus stricticollis Jeannel, 1948
  * Harpalus subaeneus Boheman, 1848
  * Harpalus subcylindricus Dejean, 1829
  * Harpalus subphaedrus Basilewsky, 1946
  * Harpalus subtruncatus Chaudoir, 1846
  * Harpalus suensoni Kataev, 1997
  * Harpalus sulphuripes Germar, 1824
  * Harpalus sundaicus Schauberger, 1933
  * Harpalus sushenicus Kataev, 1990
  * Harpalus suturangulus Reitter, 1887
  * Harpalus szalliesi Kataev & Wrase, 1995
  * Harpalus taciturnus Dejean, 1829
  * Harpalus tadorcus Ball, 1972
  * Harpalus tangutorum Kataev, 1993
  * Harpalus tardus Panzer, 1796
  * Harpalus tarsalis Mannenheim, 1825
  * Harpalus tenebrosus Dejean, 1829
  * Harpalus terrestris Motschulsky, 1844
  * Harpalus texanus Casey, 1914
  * Harpalus tianschanicus Semenov, 1889
  * Harpalus tibeticus Andrewes, 1930
  * Harpalus tichonis Jakobson, 1907
  * Harpalus tinctulus Bates, 1873
  * Harpalus tiridates Reitter, 1900
  * Harpalus tithonus Reitter, 1900
  * Harpalus torosensis Jedlicka, 1961
  * Harpalus torridoides Reitter, 1900
  * Harpalus trichophorus Tschitscherine, 1897
  * Harpalus tridens A.Morawitz, 1862
  * Harpalus triseriatus A. Fleischer, 1897
  * Harpalus turcicus Jedlicka, 1958
  * Harpalus turmalinus Erichson, 1847
  * Harpalus udege Lafer, 1989
  * Harpalus uniformis Motschulsky, 1844
  * Harpalus ussuricus Mlynar, 1979
  * Harpalus ussuriensis Chaudoir, 1863
  * Harpalus vanemdeni Schauberger, 1932
  * Harpalus venator Boheman, 1848
  * Harpalus ventralis Leconte, 1848
  * Harpalus vereschaginae Kataev, 1988
  * Harpalus vernicosus Kataev & Liang, 2007
  * Harpalus viridanus Motschulsky, 1844
  * Harpalus viridicupreus Reiche, 1843
  * Harpalus vittatus Gebler, 1833
  * Harpalus wadiensis Jedlicka, 1964
  * Harpalus wohlberedti Emden & Schauberger, 1932
  * Harpalus xanthopus Gemminger & Harold, 1868
  * Harpalus xinjiangensis Huang, Hu & Sun, 1994
  * Harpalus yinchuanensis Huang, 1993
  * Harpalus zabroides Dejean, 1829
  * Harpalus zhdankoi Kataev, 1990