The use of the phrase “largely irrelevant” suggests that while advances in understanding have been made, there are still important areas of ignorance and uncertainty that may not be fully resolved. This suggests that a deeper level of understanding may be needed to fully exploit potential therapeutic advances.

In other words, while we have made great strides in developing more accurate knowledge of the brain, there is still much work to be done in unlocking its full potential and exploring its many potential implications.
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