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Binary options tradingview indicators of diabetes
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With the internet as our guide, it seems no job is too large for our capable hands. Almost every kind of repair or renovation is explained in full online, many times with step-by-step video to go along with it. And in a time when every penny counts, paying a professional could mean dipping into your savings. However, attempting certain repairs can be dangerous to your house and harmful to your. There are jobs that you are better off leaving to a professional.
Windows and door repairs
Rashid Senyonjo, an engineer with HIL-Consult, cautions against carrying out repairs on windows and main doors because these are major parts of your home's security. For example, he says, in case the a door's hinge breaks or the padlock develops issues, it is always better to call an expert to rectify the problem in order to avoid problems such as being locked in or out.
He says replacing a broken window glass may appear simple but lining it up in the right position is hard if you are not a professional. He also says you might end up buying the wrong replacement materials due to lack of knowledge and experience.
Breakage in the drainage system
When she found her tap in the sink leaking, Christine Atuhaire tried to fix it herself. However, a simple twist resulted in the whole tap falling off leading to flooding in her house.
Sarah Katongole, a plumber at Izzona enterprises, says: "It is wrong to think that the solution to most drainage problems is just under the sink, as the saying goes. You will either need to go down to the P-trap to find the real cause. When you try to carry out maintenance work yourself, you may end up causing more damage although sometimes it may not be immediately noticeable."
Electrical repairs
ROGERS EGWAG: an electrician, says any projects involving electricity should be approached with extreme caution. You may be able to pull off minor repairs such as changing a light switch or installing a ceiling fan as long as you make sure that your power is turned off before you start. You can also test the switch against the breaker to be positive that there is not a current, or you can just turn off your master switch to be sure.
"When it comes to sort sackets, you will need to trace out for the cause which can only be done by a professional electric engineer," he warns.
Samuel Kizito, an electric engineer, says: "Extending or replacing circuits is dangerous if you do not know what you are doing. One wrong move could burn your house down, and a shock could result in injury or death.''
Roof works
Repairing a roof is not recommended for a do-it-yourselfer for one reason – it is easy to fall off the roof. Repairing a roof shingle or two is not the toughest job in the world, but getting up and down and carrying your tools with you poses the risk of injury or death. It is also tiring work, and when you are tired, you are more prone to make a mistake.
Senyonjo stresses that roofing repairs will require proper measurements or else one will mess up the entire structure.
During the contraction process or renovation, a professional engineer is equipped with all the safety gadgets such as gumboots, overalls, gloves and a head lamp.
Whether we like to admit it or not, there are home repairs best left to professionals no matter how many home renovation shows we watch. So, grab your phone instead of a hammer the next time you are faced with a large or potentially dangerous renovation.
Gas appliance repairs
According to online portal The Freeman.com, a typical home may have several different appliances that run on gas such as the oven and hot water heater.
Some homeowners feel like a hot water heater replacement is within the realm of their capabilities, and this is when accidents happen. Like water, gas will always find a leak. So while you may have done a good job in cutting off the gas supply line and moving the stove, you may not have been as careful when hooking it back up.
The end result of what you thought was a simple fix could lead to accidental carbon monoxide poisoning - something that has already killed many people.
Tiles and driveway pavers
Installing paving stones may look simple but it is a process that requires perfect positioning and measuring and even the tiniest mistake can ruin the work. "Without the knowledge, most people end up buying poor quality materials that will only last for one to three months,'' say Senyonjo.
"For instance, many people will fail to differentiate between wall, floor and bathroom tiles. That is why sometimes the bathroom is slippery leading to falling and associated injuries," he adds.

subset: commoncrawl


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Water conservation is important here in #CentralOregon. By piping the two main canals, COID's district would conserve 82,752 acre-feet of water. This significant amount of water! It's equal to an increased flow of 276 cfs in the Upper Deschutes River during the winter months. Read more about our modernization plan and how it will better help us to responsibly manage our water supplies. http://bit.ly/2ttR5xYRead more
Five years ago, you didn't see permanent organic produce sections in stores like Safeway and Fred Meyer's – now you do. Learn about two local farmers who teamed up to continue to fight for the positive trend to make locally grown, organic produce available and affordable to consumers on a broader scale. http://bit.ly/2u3kj3LRead more
Giveaway time! We're giving away 2 pairs of concert tickets/passes for the Deschutes County Fair & Expo Center coming up, Aug. 2nd – Aug 6th. How to enter? 1. Tag a friend you'd like to go to the fair/concert with in the comments below! 2. Make sure you have Liked our page. The contest will end on Monday, July 31st and the winner will be chosen at random, and announced at noon! Each winner will win a pair of tickets to one of these two concerts at random, as well as 2 fair passes: MONTGOMERY GENTRY, Wednesday, August 2nd M.E.LIVE/MELISSA ETHERIDGE, Thursday, August 3rd The Deschutes County Fair is a huge supporter of local agriculture, farmers, ranchers, and other businesses – we couldn't be more excited to support them!Read more
#Agriculture supports our local #farmers & #ranchers, this we know. It also supports countless retailers, service providers, firefighters, police officers, doctors, teachers and business owners in Central Oregon Find out how what's good for agriculture is good for our local area, and how you personally help protect our most precious resource: http://bit.ly/23ybhY7Read more
Don't let summer months slip by without taking advantage of locally grown organic produce! Brook's Alley in downtown Bend is open every Wednesday 3-7pm thru October 11th. Friday's market at Mtn. View High School is open Fridays 2-6pm and runs thru August 18th. You'll feel good inside because you supported #LocalAgriculture – and your tummy will thank you.Read more
Big news! Tumalo, Swalley, and Central Oregon Irrigation districts will be applying for a portion of recently appropriated funding for water conservation projects in the region. These projects will improve efficient use of water within the districts while allowing the districts to dedicate conserved water instream. It's important for the community to voice their support of this process before the cutoff TODAY! Please watch the video and follow the link to send your email of support for our river and our community – and a BIG thanks to the Deschutes River Conservancy for supporting the districts! https://vimeo.com/226211417Read more


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The Project for Procuring Surgical and Emergency Surgical Equipment was signed over to Pohnpei State Hospital on September 4, 2014 through Japan's Grant Assistance for Grassroots Human Security Projects and the ceremony was held at Pohnpei State Hospital. Pohnpei State will benefit greatly through this project by upgrading the quality of health care, increasing the number of outpatients and inpatients. This project will provide $89, 475 for the Hospital.
The hospital hopes that in receiving these equipment, Pohnpei State will have fulfilled its role as the principal referral Hospital and improve its performance as a leading hospital in the FSM. The project also aims to replace the present broken/obsolete medical equipment.
Ambassador Masaki Sakai, presented his remarks hoping that granted equipment contributes to enhancing medical service and hope the equipment's will serve to make these objectives a reality. Ambassador also reaffirms the commitment to contributing to foster and maintain the strong bonds of friendship through the improvement of healthcare in the FSM and other sectors. Director of Department of Health Services, Pohnpei StateMr.Paulino Rosalio acknowledged Ambassador for the generous grant and ensured that they would treat equipments properly and utilize to promote more effective use of them properly.
Present at the ceremony were staff of the Hospital and the Embassy of Japan.
Japan Grants Surgical Equipment to Pohnpei State Hospital
September 8, 2014


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In-store items and prices at Fry's retail locations may vary.
Copyright Fry's Electronics, Incorporated.M prices good for online purchases only.Some online offers are subject to prior sale and may not be available.Fry's Electronics, m and m are registered trademarks of Fry's Electronics, Incorporated.Original purchase receipt (2) Fully-completed rebate form (3).Jul 21, 2016, frys, electronics has many co-branded rebates running at all times.These are the ones.Frys is involved in and provides a Rebate Receipt and website for.If you are sending away for a manufacturer s rebate,.Original purchase receipt (2) Fully-completed rebate.Please contact our customer service.I sent in the.291 reviews Deceptive Cadence (The Conor McBride Series - Mystery Suspense Thriller Book 1) (The Virtuosic Spy) Winner, 2014 Finalist Medal for Best Action/Adventure Novel, Next Generation Indie Book Awards Readers who love thrillers with heart will find everything theyre looking for in this award-winning.
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12/31/2017 CST See sale Save On NFL Hats Sale Shop By Team At NFL Shop Visit the official store of the NFL.(5 days ago) Here are the winners for our recent giveaway: Family 4-Pack of Tickets to Magical Winter Lights Leona ( email protected ) Liz ( email protected ) Stephanie.12, 2017 6:48.(14 days ago) Discount Coupon Code: Magical Nights of Lights at Lake Lanier Islands #magicalwinterlights - Instagram photos and videos.198 reviews Untouched (The Girl in the Box Book 2) Still haunted by her last encounter with Wolfe and searching for her mother, Sienna Nealon must put aside her personal struggles when a new threat emerges Aleksandr Gavrikov, a metahuman so powerful, he could destroy.3D Mode recommended for Ages 7 Includes download code for Super Mario Kart (Super NES).4 verified offers for December, 2017.35.00 was.00, save 20 On Beauty Products.(21 days ago chuck invites you to see millions of twinkling lights this winter at the Magical Nights of Lights at Lake Lanier Islands Resort, starting November 16th through.
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Peer reviewed
ERIC Number: EJ605091
Record Type: Journal
Publication Date: 2000
Pages: N/A
Abstractor: N/A
Reference Count: N/A
Testing the 4-M Model: Introduction.
Myers-Scotton, Carol; Jake, Janice L.
International Journal of Bilingualism, v4 n1 p1-8 Mar 2000
Introduces this special issue of the journal and reports on new directions in the research that began with code switching, but increasingly has come to include other types of language contact phenomena. The articles in the issue report on analyses and explanations for a variety of outcomes in bilingual production. (Author/VWL)
Publication Type: Journal Articles; Reports - Research
Education Level: N/A
Audience: N/A
Language: English
Sponsor: N/A
Authoring Institution: N/A


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Gave this place a try after spending the day at the lake. Burgers are tasty, and fries are good. The milkshakes are a hit with all, they even do malts. Pretty good place, the family liked it.
Own or manage this property? Claim your listing for free to respond to reviews, update your profile and much more.


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Canon 99 Error Code
Remedy Change the print data or print settings, and try printing again. What are the lenses that work without problems? To prevent that from happening, which could damage the component either from heat or overstressing a mechanical part the camera shuts down & gives the error code. Remedy 1 Check the scan settings, and try scanning again. http://onewebglobal.com/error-99/canon-e99-error-code.php
View the answer I have this problem too Subscribed to new answers Is this a good question? Eric Rossi 40.590 weergaven 7:34 Canon Eos Error 99 30D - Duur: 4:28. What does that mean? Me too Report Inappropriate Content Message 1 of 3 (13,708 Views) Reply 1 Kudo Tim Product Expert Posts: 389 Registered: 10-17-2012 Re: Error 99 Options Mark as New Bookmark Subscribe Subscribe http://improvephotography.com/10372/error-99-canon-camera-fix-repair/
Canon 40d Error Code 99
Now that works too. Remedy Try delivering the device information again after the screen is cleared from the client machine. (See [Device Information Delivery Settings] in "Settings/Registration Table.") #851 Cause 1 There is insufficient memory Try using a different card. Did all that then called Canon.
Cancel Add your answer bailux will be eternally grateful. Copyright 2016 Improve Photography LLC. I get err99 on and off with my very old 350D. Canon Eos Error 99 This should help you to narrow the issue to either the lens or to the camera.
Cancel Art Hasku Rep: 1 Posted: 07/07/2014 Options Permalink History What If It Works With One Lens And with the 2 others dont work? Canon 99 Error Message Reading mode: Light Dark Login | Register NewsReviewsArticlesBuying GuideSample ImagesVideosCamerasLensesPhonesPrintersForumsGalleriesChallenges Forum indexCanon Rebel (EOS 300D-760D) TalkChange forum Error 99 Fixed Started Jul 20, 2004 | Discussions thread Shop cameras & lenses How do you get firmware onto a CF Card?
August 30, 2016 Reply Chris Big difference between a Nikon Pro film camera with no electronics and a modern digital eh?
Does that mean it is a lens problem instead? (I don't have another lens to try on the camera, unfortunately.) Thanks! Canon Lens Error 99 If I remove and reinsert the battery and do the same with the lenses, the camera is sometimes ok for a few shots but then freezes up again. I had a similar problem with a ids I bought also from eBay and "inspired" by your input, I solved it after connect it to the AC charger instead of the We tried #4 but failed to find the battery on the 1000D.
Canon 99 Error Message
More specifically, follow these steps to perform a full power cycle: (1) Turn off the camera, (2) Remove the lens and put on a body cap, (3) Remove the memory card, http://www.imagerunneradvance.com/c5000_manual/contents/trb_004b/func.html It can also be a lens problem or flash problem. Canon 40d Error Code 99 Thanks for your suggestions.Bob Atkins , Jul 25, 2015; 12:01 a.m. Canon Slr Error 99 Thank you, great article.
may i get help July 30, 2015 Reply Jure Kovac Nothing worked for my eos30d, but anyways great post. check over here It was not able to communicate with the camera. April 2, 2014 Reply RDW I had this infamous error on my Canon 5DII and had to send it off to Canon for repair. If you see the "ERR 99′ with a different Canon lens attached, then the camera should be serviced. Canon 50d Error 99
Could be a shutter issue or some kind of memory issue or one of a dozen other faults. Then I got their response - -$433 to clean and repair. Remedy Contact your System Manager. #889 Cause You cannot scan or print the document because it has Scan Lock Information embedded. http://onewebglobal.com/error-99/canon-error-code-99-40d.php Err99 means that electronic circuitry in the lens has failed.
Powershot G1x, EOS 1Ds Mk III, EOS 1D Mk IV, EF 8-15mm f/4L USM Lens, EF 16-35mm f/2.8L II USM,EF 24-70mm f/2.8L II USM, EF 70-200mm f/2.8L IS II USM,Sigma 150-600mm Canon Error 99 Shutter If you do not know the password, contact your System Manager. #849 Cause Device information could not be delivered because the client machine is processing a job. Turn off the camera, remove lens, remove battery, remove cf.
January 17, 2016 Reply Patricia This problem started from the very beginning since I got my Rebel Xti way back when (!), and never bothered to ask anyone why.
Budric Jul 20, 2004 How many pics have you taken?? If you've cleaned them and you still have the problem, it could be electrical deeper in the camera. When I change the mode to A-DEP or further, the Err 99 appears again. Canon Error 99 Repair Cost I fixed this by updating the firmware to 1.0.7 available from Canon Japan .
Cause 2 An error occurred because the main power switch was turned OFF while a job was being processed. I have two cards and this is the 1gb that came with the camera. Inloggen Delen Meer Rapporteren Wil je een melding indienen over de video? weblink Put the other lens, in place of the lens in question, on the camera body.
I don't have this problem when using any of my other Canon lenses. Marcus - my other Canon lenses that don't have this problem on the 300D are: 10-22, 70-210 f/4 IS, 100mm macro and 400 f/5.6.Louis Solomon , Jul 24, 2015; 08:44 a.m. September 12, 2016 Reply F. If you found this post to be helpful, please subscribe to my FREE photo tips newsletter and I'll email you with my very best blog posts and photo tips. Click here.
Ryan Mack 12.032 weergaven 1:32 Problem Canon 17-85 IS - Err01 Err99 - Duur: 4:16. Also the battery must be out of the camera for a long time, a few hours or so."Did not update the firmware since I bought the camera in 2009."<-----This is not http://web.canon.jp/imaging/eosd/firm-e/... I just started messing with it , and now I am able to get it to work without the error 99 when I have the lens zoomed out further than 25
Cause 3 The spool memory is full when Use Spooler in Network Settings in Preferences (from the Settings/Registration screen) is set to 'On', therefore, data received from the host could not The original lens with it 18-55 don't work any more err99 again. All OK! October 2, 2014 Reply Gerry O'Donnell Hi, I may have found a solution.
After going to the local repair place, who told me it would be $300 to replace the circuit board, I discovered that I can take unfocused pictures with no error with Remedy Contact your local authorized Canon dealer. #856 Cause The job was cancelled because there was not enough free space on the hard disk to store the temporary data. Over Pers Auteursrecht Videomakers Adverteren Ontwikkelaars +YouTube Voorwaarden Privacy Beleid & veiligheid Feedback verzenden Probeer iets nieuws! Was this answer helpful?
Toevoegen aan Wil je hier later nog een keer naar kijken? Main battery out and back - no good. Insert a fully charged battery, and turn on the camera. 5.


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Not known Details About chiropractor north hollywood
Your Over-all wellness may be resolved by a nutritional program made especially for you, making use of full foods supplementation and herbal nutritional supplements, and dietary recommendations. The particular procedure is easy and immediate, with your body giving all of the information and opinions needed."
Sending an e-mail working with this web page isn't going to warranty that the receiver will receive, go through or reply to your email.
Precisely what is chiropractic cure? Chiropractic cure is another medicine to treat back soreness by manipulation of your spine. Chiropractic technique entails use in the practitioners' palms to push, pull and if not adjust the spine, neck, joints and encompassing delicate tissues.
Like all healthier routines, each particular person need to determine for them selves how deep their motivation to their health and fitness probable is. Although we can endorse what we predict could be most effective in your circumstance, we respect your belief about how A lot of the precedence your chiropractic care plus your overall health will probably be. Without force, we respect your decisions about just the amount reward is right for you.
The brand name names, logos, visuals and texts would be the residence of these 3rd events as well as their respective proprietors. Should you have any concerns or tips regarding this subject, you are welcome to Speak to our customer assist team.
In particularly rare circumstances, manipulation with the neck has damaged blood vessels or brought on strokes. The screening approach, having said that, is made to detect men and women at superior risk.
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Chiropractic exploration continues to doc the relationship concerning a healthful backbone, healthier nervous technique perform and somebody's optimum health and fitness prospective.
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Hi there, this photo is about Catskill's Empire Work Center – Butcher Block Island, Pull-Out Leaves (ordinary Kitchen Movable Island #3). It is a image/jpeg and the resolution of this image is 728 x 438. This post's file size is just 40 KB. If You want to download It to Your computer, you might Click here. You also also download more attachments by clicking the picture below or see more at here: Kitchen Movable Island.
5 photos of Catskill's Empire Work Center – Butcher Block Island, Pull-Out Leaves (ordinary Kitchen Movable Island #3)
The thing you should contemplate will be to set a great budget, typically, the buying price of kitchen cupboards is about half of the general budget for that home. Decide on a store or a company that is trustworthy and supply warranty time. Then came alone to choose the quality of at this stage you have to know that choosing cupboards with high-quality wood content can be a lifetime expense, lumber and also other materials.
Consequently select the right lumber resources that give top and appearance quality despite the value is slightly more costly. Choose colors and finishes that you want to your kitchen units, in case you guide Catskill's Empire Work Center – Butcher Block Island, Pull-Out Leaves (ordinary Kitchen Movable Island #3) on makers, be sure you fit your individual feel. You'll be able to select the shade of white, dark, or brown in completing shiny, dull or flat finish. Pick a style to accommodate you or remain in the entire layout of the property, you'll be able to select the style of place (rural), contemporary or traditional-style.
At this time there happen to be kinds and different types of Kitchen Movable Island which might be bought soon the market. Nevertheless, when the units inside the kitchen within the form so that has been on the market do not fit your requirements, guide oneself from builders or the suppliers would be the simplest way. Just be sure to pay for focus on the budget that you simply have made. You're able to pick units while in the home that may be constructed to cut back the budget if you learn a budget exceeds the restriction.
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Diverge Cropped Skinny Jeans
Diverge Cropped Skinny Jeans
Mid rise cropped skinny jeans with heavy distressing down the front. Traditional 5 pockets with button and zip fly closure. Little stretch in denim.
*Machine Wash Cold
*98% Cotton 2% Spandex
*35.5"/90 cm Top to Bottom 25.5"/65 cm Inseam - Measured from Size 1 (Model is 5'7/ Wearing a Size 1)
*Refer to Bottoms Size Chart #2


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So, the solution that's predicted from you must be something similar to," As straightforward as it can sound, this problem is truly an exact tough one!
Prose isn't only reading. Ordinarily, all stanzas are comprised of equivalent variety of outlines in one poetry. Studying a sentence may increase the drama. To put it differently, don't embellish. Also, I didn't care for the songs that patterns through the entire movie. Presently the other components of poetry are involved. This was another contrived part of film. That is my primary writing essay complaint of the variant of Gatsby.
There are a few test issues:sort "a" query which includes 1 mark:1.
He was fascinated within the point of light of the author contrary to the plots. This article isn't heading to claim culpability or level of culpability of numerous components of the German citizenry. That is, utilize an opener which can be associated to your own essay topic. As a result of this, I'd suppose, I'm doing this investigation. The writer desired to struggle for education. I'm creating this as a kid. Why, kill him, clearly, visitors mentioned. He was the 1st author in history to be a big celebrity. A enormous literature was written on this particular problem Possibly a small should to be mentioned about spritiual poverty.
Where the problem set, it very quickly became apparent.
That is the reason poverty is a superb educator. Physical poetry, within this poem he's indicated his attitude towards love. Rhyme aids give structure to all of the themes a poet desires to cover in a specific peace. I will be capable to aid visitors fully delight in an adequate poetry, along with I may assist future poets radically enhance their craft, but I actually don't promise to become a poet myself. I was quite impressed by his wisdom. I was already abreast with the narrative. This element isn't employed in every poem.
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Just among the verses about death. You need to be powerful and packed with guts. Slowly knowledge started to dawn on him. That innovative accomplishment, however, isn't however the creating of the poetry. Therefore that, an excessive amount of mind of the process for creating or talking can get within the manner of what one is attempting to attain, with regards to suppressing Casual recognition. He stayed upbeat because of effective book sales. Do not make the major mistake of studying the complete Darwin Awards publication by signifies of your child before they begin studying their very own novel.
Personal parts wont stand out anymore if thereis a lot of there.
There's that one factor about a crucial novel. Once agan permit me to say it isn't a regular book evaluation. He's understood among the best detective and science fiction writer. Some consider Jones Ince was really shot within the middle of his own forehead, and didn't expire of the heart attack as medical care examiner reported. Ellroy's novels have a tendency to be instead'visual' for the reason that method. Today we all understand it really isn't pleasant to chuckle at some one otherwise's misfortune, but several of these stories are hilarious. When composing an everyday lifestyle storyline, you are able to skip about and play with numerous stories in your entire life. This story is jammed full of moral lessons. Don't worry, I'm not likely to assess every single narrative.
It truly is not bad to apply making use of your new terms.
Some people desire setting aside a particular time with this sort of imaginative writing varying between five moments and maybe even complete hrs. There's no tactic to educate tradition within the identical manner that vocabulary is taught. By the time which I attained Middle School, I'd learned the training. I really consider as I'm writing this it'll aid some one in their own prayer life with Lord.


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Related News:
NASA to fly helicopter on the 'red planet': 2020 missionThe rotor craft weighs in at only 1.8 kilograms, and it will fly to Mars attached to the underbelly of the rover. In a press release, NASA said the Mars Helicopter is "considered a high-risk, high-reward project".
Saudis will seek nuclear weapon if Iran does, minister tells CNNIranian President Hassan Rouhani has argued that his country can seek to upscale their nuclear arsenal "without limit" now. A journalist asked him a question during an interview that will Saudi Arabia develop a nuclear bomb itself.
Gunman shoots, injures Pakistan inside ministerThe row sparked a three-week sit-in last November by a previously little-known Islamist group, which paralysed the capital. The gunman, the attacker was arrested, his identity and the motive of the crime yet no information is available.
Cancer Patient to Attend Capitals vs. Lightning Game 3 in Between TreatmentsAdding linemate Evgeny Kuznetsov's career-high 12 shots on goal, the duo accounted for more than half of Washington's 38 shots. Road teams that have taken a 2-0 series lead in the National Hockey League conference finals have an 18-1 series record.
Wayne Rooney is last of the 'street footballers' - David MoyesWe've not been in any hard situations because we're two adults and we talk'. Until someone tells me I'm no longer wanted then I have a contract.
Celtics' Stevens remains humble as team continues to thriveCavaliers' defenders can get twisted around on pick-and-rolls, and particularly dribble handoffs (which the Celtics run a lot of). Philadelphia's Joel Embiid missed a potential game-tying layup with 12.5 seconds left before the 76ers turned the ball over.
U.S. unilateral withdrawal from JCPOA, unethical: Pres. RouhaniThe head of the elite Revolutionary Guards also warned against relying on foreign powers. On Wednesday, Jafari cast doubt on European nations' ability to save the accord.
Walmart Eyes India E-Commerce Potential with Flipkart AcquisitionFlipkart is India's largest e-commerce group on the basis of sales but has been fighting off a huge challenge from Amazon. For the US retailer, acquiring a stake in Flipkart enables it to tap into India's retail market without building stores.
Red Sox, Yankees To Face Off In London In 2019London Stadium , the site of the 2012 Olympic Games, which will take on a baseball configuration for the event. The arrival of the game in London follows talks with Sadiq Khan when he became mayor in 2016.
Jada regrets dating Will Smith!Will and Jada are Hollywood legends, Jaden is becoming an icon in his own right, and Willow's musical career is still on the rise.
Selena Gomez is Focused on Making Hits & Recording Her Third Studio AlbumIt was all in fun, of course, and Gomez played along when she finally broke her social media silence on Wednesday morning. It shows a video of her in her dress with her back to the camera, sprinting towards her ride.
Rajasthan Royals elect to bat and leave out D'Arcy ShortOn the other hand, Punjab have great batsmen like Gayle, Mayank Agarwal, Karun Nair and Marcus Stoinis . Despite Rahul fetched 32 runs from the last 12 deliveries, Rajasthan were still 15 runs short.


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Home-Health and Hospice Care -- United States, 1992
CreatorUnknown creator
Bibliographic Citation
MORBIDITY AND MORTALITY WEEKLY REPORT 1993 October 29; 42(42): 820-824
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An Act Relating to Individual and Group Accident and Health Insurance Coverage for Home and Inpatient Hospice Care and Making Penalties Applicable Unknown creator (Iowa. Legislature. House, 1973-02-23)
Caring to the End: Policy Suggestions and Ethics Education for Hospice and Home Health Care Agencies Drane, James F. (1997)
RECOMBINANT DNA RESEARCH, VOLUME 16: DOCUMENTS RELATING TO NIH GUIDELINES FOR RESEARCH INVOLVING RECOMBINANT DNA MOLECULES, JULY 1992-DECEMBER 1992 National Institutes of Health [NIH] (United States) (1994)


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Mediteranean Cruise?
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Mediteranean Cruise?
I am taking a cruise in the mediteranean in october on the Costa Riviera. I understand everything on board is very expensive. Does anyone have any suggestions as to what if anything I could bring from home that would save me some money?
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Almost everything is included on your cost of the cruise. The only possible thing I can think of to bring might be liquor if you drink and don't want to spend the money for cocktails. Your off-shore excursions will be an extra cost and you should get all that info. before you leave on you trip.
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In response to "Death Watch: Easley's 20th execution pending this week," by Patrick O'Neill (May 4), I would to say that while there are serious moral arguments to be made against the death penalty, one must remember that some criminals simply deserve to be killed for the crimes they commit. Period. Any adult who murders children deserves no less. The argument that clemency should be granted in such cases is an insult to the intelligence of the living and a sacrilege to the memory of the dead.
Christopher Yahnovich
OK, he's a wingnut
First off, thanks for the info on the Kansas invasion that took place in Durham this week: aka the visit of Mr. Phelps to the Bull City to protest the Durham School of the Arts production of The Laramie Project ("Turn your back on him," May 4). Good job.
But you folks must be crazy, calling this extremist, this terrorist, this wingnut a Reverend. PUH-LEEEZE. Phelps is no reverend regardless of what he claims. The Indy shouldn't be glorifying hatemongering masquerading as religion in this fashion. An extremist Christian is just as destructive as an extremist Muslim.
There are legitimate conservative Christian leaders. Phelps is not one of them. We need to start making distinctions, and say in no uncertain terms, what is simply beyond the pale. The place to start is calling a wingnut a wingnut.
Jim Senter
In a story in the April 27 issue about a bill in the General Assembly to allow higher alcohol content in beer ("Beer bill coming to a head"), the name of Giorgios Hospitality Group was misspelled as was the name of the firm's beverage director, Ken Rosati. Giorgios Hospitality Group runs five restaurants, four in Durham and one in Chapel Hill.
The author for reviews of new releases--the compilation Heard Together and The Rosebuds Unwind EP--was misidentified in last week's music section. The reviews were written by Chris Parker.
talk back.
Send no more than 300 words to firstname.lastname@example.com; to P.O. Box 2690, Durham 27715; or fax 286-4274. Include name, phone number and mailing address for verification.


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FOR THE FIRST TIME this term, the Supreme Court sat yesterday with a full complement of nine justices. And one announcement made by the court means that the newest member, Justice Anthony Kennedy, has arrived just in time for an extremely important case: the court agreed to decide the constitutionality of the Ethics in Government Act, which authorizes the appointment of special prosecutors in cases involving high government officials. As an indication of the urgency of settling this question, the court will hear the case on an expedited basis -- arguments have been set for April 26 -- and will hand down a decision before the term ends in June.
This prompt action is welcome. The special prosecutor law was passed in 1978, but until recently no investigation had produced an indictment. Now, Michael Deaver and Lyn Nofziger have been convicted in prosecutions brought by special prosecutors, and two minor figures in the Iran-contra scandal have entered guilty pleas. It has been clear for some time that a serious challenge to the law would be made and would have to be resolved by the Supreme Court. Last year, most of the special prosecutors who were then conducting investigations accepted parallel appointments in the Justice Department in order to preserve their work should their authority under the ethics law be invalidated. Two did not. Whitney North Seymour, who prosecuted Michael Deaver, and Alexia Morrison, now investigating three former Justice Department officials -- Theodore Olson, Edward Schmultz and Carol Dinkins -- chose to continue without this backup authority. The case before the Supreme Court involves the Olson inquiry, but the ruling could presumably affect the Deaver conviction as well.
The U.S. Court of Appeals for the District of Columbia has ruled the special prosecutor law unconstitutional, but the question would have gone to the Supreme Court no matter what the finding below had been. The appellate court was divided 2 to 1, and strong opinions were written on each side. Does the ethics law, as the majority found, depart from the constitutional framework of balance of power in authorizing an independent, court-appointed attorney to prosecute crimes? Or, as the judge in dissent and the majority in Congress believe, is it the only practical mechanism available for dealing with wrongdoing at the highest levels of government? There will be a final answer before summer.


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Articles from the March 21, 2012 edition
Sorted by date Results 1 - 25 of 32
Basket factory
Genny Rockwell shops for Easter baskets for her almost-4-year-old twins at the Grand Coulee Dam Senior Center on Main Street Monday. The seniors made 365 baskets this year, and are selling them for... Full story
Tribal members hear terms of settlement
The only thing certain about how a $193 million settlement with the U.S. government will be used is that 20 percent of it will be paid out to members of the Confederated Tribes of the Colville Reservation. Exactly how and when the $38.6 million in... Full story
Crime watch idea raised
Two Electric City council members concerned about the amount of criminal activity are interested in developing a "neighborhood watch." Lonna Bussert and Birdie Hensley discussed their interest at last Tuesday's council meeting, and have asked... Full story
News Briefs
TV program on dam set for April Amercian Experience will air a new documentary on Grand Coulee Dam April 3. It will be carried on Spokane's KSPS-TV public television station at 7 p.m. A repeat broadcast is scheduled for April 7 at 2 p.m. "Grand... Full story
Town will put $30,000 to school upgrade
The town council of Coulee Dam has voted $30,000 worth of energy credits to help the school district upgrade Lake Roosevelt High School's gymnasium. The council heard of a request by school district Superintendent Dennis Carlson and Jayson Schmidt... Full story
Famous bull riders plan to attend event
Rodeo fans will want to take advantage of the Ridge Rider's Banquet and Auction April 2, and get to meet some of the world's premium bull riders, including the world champion from Grand Coulee, Shane Proctor. The banquet and auction is a new... Full story
Port district hires help for golf course
Port District officials have taken the first step in getting Banks Lake Golf Course open. District officials hired George Davis III, as grounds and greens keeper. Davis had worked the past year in that capacity, but was released when the operators... Full story
Crest Drive improvements funded
by Roger S. Lucas The town of Coulee Dam has received a Washington State Department of Transportation grant for $302,750, for major improvements to Lower Crest Drive, the council learned at its meeting last Wednesday night. Public Works Director... Full story
Elmer City water extension team earns award
The creation of a new water service for Elmer City has garnered an award for those who made it happen. Elmer City's water extension team, made up of Mayor Mary Jo Carey, Town Clerk Renée Tillman and Public Works Director "Jimmer" Tillman,... Full story
Club plans for bigger, better shred day
Grand Coulee Dam Rotary Club is planning to tear up even more stuff than it did last year in an upcoming free community shred day. The club's first free community shred day last year had been scheduled to run from 9 a.m. to 1 p.m, but community... Full story
New DUI law stiffens penalties, opens records
After a Friday night of happy-hour and late-night drinking, many fail to consider the repercussions of driving while impaired, but legislators have. Various changes to the driving-under-the-influence law have unanimously passed the Legislature and... Full story
Building a base
Workers sculpt the hillside above Coulee Dam to prepare it for new high voltage power line towers. Two sets of towers and lines will come down in process, before more are erected to hold up 18... Full story
God gives us the right to choose
There is nothing I want more than for everyone to be saved. I lie awake wondering how God will save us, and I always come back to the same answer again and again. God will save us through his love, which he revealed to us in his son Jesus Christ. Too... Full story
Community support appreciated
As we approach what would have been Dad's 85th birthday, we are reminded what a wonderful sendoff the community gave to our dad, Rod Hartman. The tears still come as we think of the fire trucks, ambulances, police cars sitting patiently by, waiting... Full story
Morgan's Musings - Morgan muses yet again
First off, I really appreciate the emails, phone calls and chit-chat from those of you in the street and on the river bank about the old Musings columns during the past decades. Believe you me, it... Full story
Coulee Recollections
Ten Years Ago The goal of flying lots of American flags in all four of the towns in the Grand Coulee Dam area might not occur after all. Members of the Elmer City Town Council voiced their concerns about the project last Thursday. OMAK - the largest... Full story
It's Not Just Congress: Citizens also have room to improve
As a member of Congress, you get used to being graded. Interest groups send you questionnaires, check your voting record, and then issue their "report cards." Editorial writers opine freely on your performance. Pollsters issue monthly updates on... Full story
Terri L. Chamberlain
Terri L. Chamberlain, 47, of Electric City, passed away suddenly Monday, March 12, 2012, at Coulee Medical Center in Grand Coulee. Terri was born in Blackfoot, Idaho, on May 4, 1964, to Rex and Lou... Full story
Harold W. Dugan
Memorial services for Harold W. Dugan will be held at noon, Saturday, March 31, at Spring Canyon Cemetery. It will be with OES ceremony and full military honors. Dugan passed away December 22, 2011.... Full story
Meetings and Notices
Chamber to Meet The Chamber of Commerce will meet at noon on Thursday, March 22, at Pepper Jack's Bar and Grille. Eagles #2577 Meetings The Eagles #2577 Aerie meets on the second and fourth Tuesday of every month. The next meeting is Tuesday,... Full story
Mullen/Turcotte have a girl
Nichole Mullen and Donald Turcotte of Omak are proud to announce the birth of their daughter Zoe Renae Turcotte on Tuesday, March 13, 2012, at Coulee Medical Center in Grand Coulee. She weighed 6 lbs., 9 oz., and was 19-1/2 inches in length at... Full story
It's a girl for the Goetzes
Jason and Beth Goetz of Odessa are proud to announce the birth of their daughter Rylee Marie Goetz on Thursday, March 15, 2012, at Coulee Medical Center in Grand Coulee. She weighed 9 lbs., 2.4 oz., and was 21.5 inches in length at birth. Siblings... Full story
Native speaker to bring message to Coulee Dam church
An American Indian evangelist, singer and songwriter will minister at the Coulee Dam Community Church Sunday, April 1, at 11 a.m. Morning Sun Yellow Pony, from her home in Montana, has been traveling for eight years with her "Healing Wings Ministri... Full story
Emerson to exhibit cattle at regional show
Katie Emerson, of Grand Coulee, will exhibit Angus cattle at the 2012 Western Regional Junior Angus Show in Reno, Nev., April 21-23, reports Robin Ruff, director of junior programs of the American Angus Association. Emerson, a junior member of the... Full story
The reigning McClures
Dave McClure, left, and Andrew McClure, father and son, accept the adulation of the throng at Lake Roosevelt High School's dual pageants for Mr. GCD and Mr. LR March 13. — Emily Williams... Full story


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Manufacturer Part Code: BOXNUC7I3BNK
Order Code: PBINT-NUC7I3BNK
Category: Intel NUC Devices
Intel NUC BOXNUC7I3BNK i3 7100U 2.4GHz Baby Canyon Barebone Slim Kit
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Intel NUC
CPU Included
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Number of Cores
Dual Core
Number of Threads
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i3 7100U
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Latest News on Balrampur
Showing of 0 - 10 from 12 results
Balrampur - Total results - 12
May 30, 2018
Youth BJP Dalit leader found hanging from tree in West Bengal, chilling message on body says he was 'punished'A young BJP Dalit leader was found hanging from a tree on Wednesday in Purulia district's Balrampur.
Apr 24, 2018
Eight-year-old girl raped at knifepoint in Uttar Pradesh's Balrampur, accused arrestedAn eight-year-old girl was allegedly raped at knifepoint by a man in a village in Uttar Pradesh's Balrampur district, police said on Tuesday.
Nov 16, 2017
IED blast along Chhattisgarh-Jharkhand border kills seven security personnel; Naxals triggered violenceSeven security personnel were injured when Naxals allegedly triggered an IED blast and fired at them along the Jharkhand-Chhattisgarh border on Thursday, the Jharkhand police said.
Aug 26, 2017
Uttar Pradesh floods: 25 lakh people affected by floods, says state govtMore than 25 lakh people living in 25 districts of Uttar Pradesh have been affected by floods this year, the state government said on Saturday.
Aug 20, 2017
Uttar Pradesh floods: Army evacuates people marooned in Gorakhpur, 250 civilians rescued in 24 hoursThe army on Sunday rescued people marooned in flood-hit Gorakhpur city in Uttar Pradesh, a military official said.
Aug 20, 2017
Uttar Pradesh floods: 69 dead, over 20 lakh affected in 24 districts as rains continue to lash stateWith fresh flood-related deaths reported from across Uttar Pradesh, the death toll in the current wave of floods has risen to 69.
Aug 18, 2017
Uttar Pradesh floods: Indian Army steps in for rescue operations; Yogi Adityanath conducts aerial surveyThe current spell of floods in Uttar Pradesh has claimed 36 lives, prompting authorities to seek the army's help for relief and rescue operations, officials said on Friday.
May 05, 2016
Bus accident in Chhattisgarh: Death toll rises to 16The death toll in the bus accident in Chhattisgarh's Balrampur district has gone up to 16, with three more persons succumbing to their injuries.
May 05, 2016
Chhattisgarh: 13 killed, 53 injured after bus falls into dry rivuletAt least 13 people were killed and 53 others injured when a bus fell off a small bridge into a dry rivulet in Chhattisgarh's Balrampur district.
Mar 07, 2016
Man in UP shot dead by MLA's brother for overtaking his vehicleA man in Uttar Pradesh was allegedly shot dead by a Samajwadi Party (SP) MLA's brother on Saturday night for overtaking his vehicle.


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Understanding how species have replaced each other in the past is important to predicting future species turnover. While past species replacement is difficult to detect after the fact, the process may be inferred from present-day distribution patterns. Species with abutting ranges sometimes show a characteristic distribution pattern, where a section of one species range is enveloped by that of the other. Such an enclave could indicate past species replacement: when a species is partly supplanted by a competitor, but a population endures locally while the invading species moves around and past it, an enclave forms. If the two species hybridize and backcross, the receding species is predicted to leave genetic traces within the expanding one under a scenario of species replacement. By screening dozens of genes in hybridizing crested newts, we uncover genetic remnants of the ancestral species, now inhabiting an enclave, in the range of the surrounding invading species. This independent genetic evidence supports the past distribution dynamics we predicted from the enclave. We suggest that enclaves provide a valuable tool in understanding historical species replacement, which is important because a major conservation concern arising from anthropogenic climate change is increased species replacement in the future.
1. Background
Species that meet in nature but geographically exclude one another have parapatric distributions [1]. While the position of parapatric range boundaries may be dynamic, their actual movement is a protracted process and has necessarily been recorded over shallow time frames only [1,2]. We here argue that the current spatial arrangement of parapatric species can be informative about distributional shifts in the more distant past. In particular, we highlight the insight provided by enclaves. An enclave is formed when part of the distribution of one member of a pair of parapatric species is isolated inside the range of the other (figure 1). An enclave could originate via colonization—analogous to a species establishing a peripatric population on an oceanic island [3,4]. Alternatively, the enclave might be a relict distribution patch of a previously broader distribution, left after incomplete species replacement—akin to rising sea levels disconnecting a continental island population from the mainland [5,6]. The likelihood of either scenario depends on the interplay between the mean and shape of a species's dispersal kernel [7] and the enclave's distance from the main distribution range [8]. We assert that enclaves of species with low vagility, relative to the distance between the enclave and the main range, provide strong a priori support for species replacement. We provide proof of concept for an enclave system involving newts.
We take advantage of the fact that contact zones for parapatric species tend to correspond to hybrid zones [1]. When, upon secondary contact, a hybrid zone is first established, and one of the species possesses a competitive edge over the other one, this would result in a non-equilibrium phase of species replacement, with the hybrid zone moving as a consequence [1,4,5]. Furthermore, for a hybrid zone initially stabilized at an environmental gradient, this equilibrium phase could be disrupted if climate change shifts the balance in favour of one of the species [9]. If the species involved show introgressive hybridization, a moving hybrid zone is predicted to leave a trail of unlinked, selectively neutral alleles in its wake, derived from the displaced species and inside the territory newly claimed by the expanding one (initially based on theoretical principles [10], later supported by simulation [11] and recently backed up by empirical findings [12]). Under this rationale, the hypothesis of enclave formation can be tested independently in hybridizing species, based on the geography of genome-wide interspecific gene flow: an enclave formed by incomplete species replacement is expected to be accompanied by a genomic footprint of hybrid zone movement, while an enclave formed by colonisation is not consistent with this pattern (figure 1).
We examine an enclave observed in crested newts (Amphibia: Triturus), located in central Serbia, on the Balkan Peninsula [13]. Part of the range of T. ivanbureschi (blue in figure 2a) is detached from the main distribution, because the range of another species, T. macedonicus (green), intervenes. The ranges of two additional species, T. cristatus (red) and T. dobrogicus (orange), border that of T. ivanbureschi in the north. Both T. ivanbureschi and T. macedonicus colonized the central Balkan Peninsula postglacially, as they expanded from discrete glacial refugia, distant from the enclave [15]. Triturus cristatus was already in place as it had a glacial refugium in the southern protrusion of its current range, where it borders T. ivanbureschi today [16]. Although T. dobrogicus expanded its range postglacially [17], it is, uniquely among crested newts, a specialized lowland species [18], and its distribution in the south is bounded by an elevational ecotone (figure 2b). Therefore, while T. cristatus or T. dobrogicus represent a biological barrier north of the enclave, it is unlikely that either species invaded the range of T. ivanbureschi over a substantial area.
As the distance between the enclave and the main range of T. ivanbureschi (approx. 83 km) is over 20 times larger than lifetime dispersal distance (approx. 3.7 km [12]), incomplete species replacement (rather than long-distance colonization) is the likely explanation for the enclave's origin. Because crested newt species ranges meet at hybrid zones that facilitate introgression [19], we are in a position to test for a genomic footprint of hybrid zone movement around the enclave. We predict that such a genomic footprint was left by T. ivanbureschi, where we assume it was superseded by T. macedonicus. We do not predict a pronounced asymmetry in introgression between T. ivanbureschi and either T. cristatus or T. dobrogicus, as we expect T. ivanbureschi's hybrid zones with these species to have been relatively stable.
2. Material and methods
(a) Sampling
We included 1–3 individuals (mean 2.645) from 251 Triturus localities; 664 individuals in total (figure 2a; electronic supplementary material, table S1). For part of these individuals the full set of sequence data (n = 308) or (mt) DNAs (n = 283) were available from previous studies [12,17,20–24], while additional individuals (n = 73) from Serbia and Bulgaria are studied here for the first time. We selected 15 reference individuals per species—three individuals from five localities positioned away from contact zones and presumed to be unaffected by interspecific gene flow—to determine whether individual markers are diagnostic (electronic supplementary material, table S1). We created a Voronoi diagram in ArcGIS 10 (www.esri.com), where each locality is represented by a Thiessen polygon that contains the area that is closer to that particular locality than to any other one (figure 2c).
(b) mtDNA sequencing and analysis
We Sanger sequenced a 658 bp mtDNA fragment for 625 individuals (following [17]) and were able to sequence a 110 bp internal fragment for 23 individuals following [12]. The clear-cut geographical distribution of mtDNA allowed us to infer mtDNA type for the 16 remaining individuals (electronic supplementary material, table S1). The 658 bp mtDNA fragments were added to the mtDNA haplotype database of [17] and collapsed into haplotypes with MacClade 4.08 [25]. To assign new haplotypes to species, we constructed a neighbour-joining phylogeny with 1000 bootstrap replicates in MEGA 5 [26], with the Pyrenean newt Calotriton asper and the marbled newt T. marmoratus (from [27]) as outgroups (electronic supplementary material, figure S1; table S2). Internal 110 bp mtDNA fragments were aligned with the set of 658 bp haplotypes and this set was then trimmed accordingly. By removing redundancy in MacClade, the internal fragments could be allocated unambiguously to species-diagnostic mtDNA type.
(c) Sequencing nuclear DNA
For all 664 individuals, we sequenced 52 nuclear markers following [22]. Details are in electronic supplementary material, text S1.
(d) Bayesian clustering analysis
We used Structure 2.3.3 [28] to estimate the fraction of ancestry for each individual derived from the four parental species based on nuclear DNA data. We used the admixture model in combination with the correlated allele frequency model with 1 000 000 iterations, after 250 000 iterations of burn-in, and ran 10 replicates. An initial Structure analysis on the set of reference individuals, in which we allowed the number of genepools k to vary from 1–20 (with the upper limit defined by the total number of localities included), confirmed k = 4 as the most likely number of genepools under Evanno's Δk criterion [29], as implemented in Clumpak [30]. Next we conducted a Structure run on the full set of individuals, in which we fixed k to 4. The lowest Q-value with which a reference individual was allocated to its respective species was Q = 0.9794. Localities allocated to two (or in some cases three) species with Q > 0.0206 (i.e. 1–0.9794) were considered genetically admixed. Structure scores (with replicates summarized in Clumpak) are presented in electronic supplementary material, table S1 and locality averages are plotted in figure 2b.
(e) Hybrid index, heterozygosity and ancestry, and test for asymmetric introgression
We determined the evolutionary origin of alleles based on our 15 reference individuals per species (see section on sampling). We found 23 nuclear DNA markers to be diagnostic, i.e. exhibiting fixed allelic differences between T. ivanbureschi (the species with the enclave) and the other three species. We determined the proportion of diagnostic T. ivanbureschi alleles present at each locality, i.e. the mean hybrid index (electronic supplementary material, table S1), and plotted this on a map (figure 2c). For pairwise species comparisons of T. ivanbureschi versus the other three Triturus species, we determined individual heterozygosity (the fraction of markers heterozygous for alleles from each parental species) and ancestry (the fraction of alleles derived from each parental species) using the R [31] package 'HIest' [32]. Locality averages (electronic supplementary material, table S3) were plotted in Statistica 7 (www.statsoft.com) (figure 3). To test if introgression between species pairs was significantly asymmetrical we determined, for each locality, whether there was introgression of nuclear DNA alleles and what the fraction of introgressed alleles was (electronic supplementary material, table S3). We used a Fisher's exact test for the presence/absence data and a one-tailed Mann–Whitney U test for the quantitative data (electronic supplementary material, table S5). To avoid the confounding effects of gene flow from other Triturus species, we excluded localities showing ancestry of additional Triturus species (based on the Structure analysis or, in the case of low frequency introgression, inferred from the genetic composition of neighbouring Thiessen polygons; electronic supplementary material, table S4) from pairwise comparisons.
(f) Geographical cline analysis
In pairwise species comparisons for T. ivanbureschi versus the other three species, excluding localities showing ancestry of additional Triturus species (electronic supplementary material, table S4), we determined the 0.5 contour for the hybrid index using the 'akima' package [33] in R. We calculated the minimum straight-line distance of each locality from that contour in ArcGIS, and gave distances for T. ivanbureschi a positive sign and distances for other species a negative sign (electronic supplementary material, table S3). For the resulting one-dimensional transect [34] we then fitted a geographical cline to the hybrid index in the R package 'HZAR' [35]. Frequency was fixed at the ends of the cline to 1 (pure T. ivanbureschi) and 0 (pure for the other species). The hybrid index is the proportion of diagnostic alleles, calculated from 23 diploid markers (minus a few missing genotypes) in one to three individuals. Therefore, it was fitted with binomial error distribution and sample size equal to the total number of genotypes (loci × individuals) per locality. Although markers are unlikely to be closely linked, they may not behave independently. Using the number of genotypes rather than the number of alleles makes our test conservative, despite this constraint. We tested five models, differing in complexity: no tail, left tail only, right tail only, mirror tails or both tails estimated separately. We used the lowest Akaike information criterion score corrected for small sample size to select the best one; in each case the score of the chosen model was at least two likelihood points lower than that of the next best fitting model (figure 4; electronic supplementary material, table S6).
3. Results
Bayesian clustering analysis with Structure based on 52 nuclear DNA markers allows us to delineate the enclave of T. ivanbureschi (figure 2b). The range of T. macedonicus penetrates north-eastwards, into the range of T. ivanbureschi and right up to the range of T. cristatus, isolating the T. ivanbureschi enclave. Several localities between the enclave and the main range of T. ivanbureschi show genetic admixture among all three species. A closer look at the 23 nuclear DNA markers that are diagnostic for T. ivanbureschi versus the three other species reveals a substantial number of T. ivanbureschi alleles in the core of the range of T. macedonicus, but not the other way around (figure 2c). Similarly, the mtDNA of T. ivanbureschi protrudes into the range of T. macedonicus (figure 2c). No such asymmetry is apparent when comparing T. ivanbureschi with T. dobrogicus, while foreign alleles reach inside the range of T. cristatus only over a short distance, just north of the enclave. Plots of heterozygosity versus ancestry support the pattern of T. ivanbureschi alleles reaching further into the range of T. macedonicus than the ranges of the other two species (figure 3).
The asymmetry in nuclear DNA introgression from T. ivanbureschi into T. macedonicus is statistically significant, based on either the presence (Fisher's exact test, p = 0.000) or the intensity (one-tailed Mann–Whitney U test, U = 1423.5; Z = −3.11; p = 0.000) of introgression. There is no such asymmetric nuclear DNA introgression into T. cristatus (p = 0.554; U = 1071.0; Z = −0.77; p = 0.341) or T. dobrogicus (p = 1.000; U = 810.5; Z = 0.47; p = 0.444). To remove the influence of a potential bias in sampling, we collapse the two-dimensional sampling into a one-dimensional transect, based on the shortest distance of localities to a 0.5 hybrid index contour. One-dimensional geographical clines, fitted to a hybrid index derived from the 23 diagnostic nuclear DNA markers for T. ivanbureschi versus each of the other three species, differ in the models that best fit the data. Only for T. macedonicus was an asymmetric cline model preferred, with a substantial introgression of T. ivanbureschi alleles into T. macedonicus, but not in the opposite direction (figure 4).
4. Discussion
Guided by the enclave observed in our crested newt system, we hypothesized a past parapatric range shift, in which T. macedonicus intersected the range of T. ivanbureschi and excised the enclave. As the two species must have hybridized in the process, we could test this scenario by exploiting the pattern of interspecific gene flow. In line with predictions for moving hybrid zones based on theory [10] and data simulation [11], we exposed a genomic footprint of hybrid zone movement, left by T. ivanbureschi, where it has been replaced by T. macedonicus. While cytonuclear incompatibilities or dispersal effects might cause asymmetry in mtDNA introgression and selection might cause asymmetry for any given locus (while not necessarily in the same direction), a genome-wide shared asymmetry in introgression over an extensive area strongly favours a scenario of species displacement with hybridization [12]. Hence, the crested newt case illustrates the predictive power of enclaves for inferring past species replacement.
Enclaves are not often reported, but, considering that three are known in Triturus newts alone (see [36,37] for two other species pairs), this does not necessarily mean that they are rare. Enclaves may well be transient, with species replacement not prevented but merely postponed locally. Indeed, all individuals from the T. ivanbureschi enclave show some degree of genetic admixture with other crested newt species. The ephemeral nature of enclaves is illustrated by a previously documented enclave (established from museum specimens) having been taken over by an invader over a period of roughly 50 years [38]. Furthermore, 'islands of alleles' derived from a displaced species that was genetically swamped out by a competitor, could be regarded as genetic fossils of a past enclave [39]. In general, being small and isolated from the main distribution makes enclave populations more susceptible to the extinction risks associated with habitat fragmentation [40].
Once an enclave is identified, it can provide crucial insights into historical biogeography. While hybrid zones have been tracked at the scale of decades [4], they are thought to stabilize quickly at environmental density troughs [10,41]. Enclaves, however, indicate hybrid zone movement over extended periods of time—an opportunity that has as yet received little-to-no attention in the hybrid zone literature [4,11]. Understanding past shifts in the mutual range boundaries of parapatric species is particularly relevant in light of the expected future increase in such shifts in response to man-made habitat alteration and climate change [9,42].
Permits were provided by the Ministry of Environment and Water of Bulgaria (permit nos 153/28.05.2008, 182/09.03.2009, 258/09.04.2010, 411/14.07.2011, 520/23.04.2013 and 656/08.12.2015). Tissue samples of museum specimens were taken from the Batrachological collection of the Institute for Biological Research 'Siniša Stanković', University of Belgrade.
Data accessibility
The data underlying this paper are archived in the Dryad Digital Repository (https://doi.org/10.5061/dryad.9k1h0) [43].
Authors' contributions
B.W., T.B., R.K.B. and J.W.A. designed the research. B.W. performed the research. B.W. wrote the paper with input from T.B., R.K.B. and J.W.A. All authors gave final approval for publication.
Competing interests
We have no competing interests.
B.W. initiated this work as a Newton International Fellow. This project has received funding from the European Union's Horizon 2020 research and innovation programme under the Marie Skłodowska-Curie grant agreement no. 655487. T.B. is a Leverhulme Fellow.
Ana Ivanović, Maja Slijepčević, Tanja Vukov, Borislav Naumov and the late Nikolay Tzankov provided samples. Pictures of crested newts in figure 2 were provided by Michael Fahrbach.
Electronic supplementary material is available online at https://dx.doi.org/10.6084/m9.figshare.c.3931906.
- Received September 8, 2017.
- Accepted October 30, 2017.
- © 2017 The Authors.
Published by the Royal Society under the terms of the Creative Commons Attribution License http://creativecommons.org/licenses/by/4.0/, which permits unrestricted use, provided the original author and source are credited.


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Etcetera: Nike, Dick's
Nike extended its sponsorship of UK Athletics through 2030, according to The Daily Mail. The existing deal, worth £15 million ($19.2 million), was set to expire in 2020.++++Dick's SG ... Log in to view full article.
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October 15, 2015
UAMS Helps Launch Plan to Improve Arkansans' Health
Oct. 15, 2015 | UAMS has joined with other state leaders in health care, education, business and government to launch a comprehensive plan to improve the health of all Arkansans.
The plan, called Healthy Active Arkansas, was released Oct. 14 at a news conference at the State Capitol attended by representatives of the different sectors that participated in developing the 10-year plan.
The plan, which can be found at www.healthyactive.org, is a culmination of almost two years of work facilitated by the Winthrop Rockefeller Institute through a leadership group that included UAMS Chancellor Dan Rahn, M.D.
A recently released Robert Wood Johnson Foundation reports said Arkansas has the highest rate of obese adults in the United States.
Gov. Asa Hutchinson announced the plan, saying Healthy Active Arkansas is a positive message for the state and is important in his goal as governor to help grow the state's economy and create jobs. He challenged citizens, businesses, health care entities and government to all get involved in putting the plan into action.
The plan contains nine focus areas tied to increasing the health of Arkansans through healthy dietary choices and increased physical activity.
The nine focus areas are:
- physical and built environment
- nutritional standards in government, institutions and the private sector
- nutritional standards in schools — early child care through college
- physical education and activity in schools — early child care through college
- healthy worksites
- access to health foods
- sugar-sweetened beverage reduction
- breastfeeding
- marketing program
Others who spoke at the news conference included Arkansas Surgeon General Greg Bledsoe; Pearl McElfish, Ph.D., research director for UAMS' northwest Arkansas campus; Michael Knox of the Arkansas Minority Health Commission; Jayme Mayo, a physician assistant representing a successful wellness program at Nabholz Construction; and Troy Wells, CEO of Baptist Health.
McElfish works with the Marshallese and Hispanic populations in northwest Arkansas, which have a higher than average rate of obesity and diabetes. She said the plan recognizes the importance of addressing environmental and other factors such as language barriers and cultural norms that affect minority populations.


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