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US, Europe & Asia Pharmaceutical Desiccant Market Analysis (Features, Segmentation) & Global Market Forecast to 2023
View Article: http://www.marketresearchfuture.com/articles/us-europe-asia-pharmaceutical-desiccant-market-analysis-features-segmentation-global-market-forecast-to-2023
Article Brief
This article provides an overview about the global Pharmaceutical desiccant market during the forecast period 2016 to 2023, there by offering insights on key present growth indicators and forecast indicators such as drivers, restrains, opportunities and trends that are involved in shaping the market.
About Desiccant
A desiccant is a hygroscopic substance which is used to induce or sustain a state of dryness in its vicinity. Desiccant are solids that absorb water and are mainly used to remove humidity that would Degrade or even destroy products sensitive to moisture
Pharmaceutical Desiccant
These types of desiccant are specially characterised for the absorption process in the pharmaceutical as well as the nutraceuticals packaging. As of now there are number of pharmaceuticals desiccant in the market. Some of the desiccants are:?
- Silica Gel?
- Activated
- CarbonClay Desiccant
- Molecular Sieves
Some of the many desiccant products that can be used in pharma desiccant packing, without causing any harm to the pharmaceutical products include desiccant canisters; humidity indicating cards, tablet form of desiccant, active vails, small sized desiccant sachets, etc. managing and controlling the level of moisture in the atmosphere is the main objective of a desiccant
Features of Pharmaceutical desiccant
- Cost effective solutions to protect drugs from moisture
- Extends drug life
- Oder control
- High absorption rate
Pharmaceutical desiccant can be used in products like
- Tablets
- API's
- Capsules
- Diagnostic kits
- And also in Nutraceutical's product packaging
Market segmentation by type of products
- Activated Alumina
- Activated Alumina balls
- Container Desiccant
- Cargo dry pack
- Chromatography Absorbent
- Aluminium Oxide Chromatography
- Silica Gel Chromatography
- Consumer Products
- Mold Away
- Storage dry
- Refresherator Dry
- Silica
- GelSilica gel blue
- Silica gel orange
- Silica gel white
- Molecular Sieve
- Molecular Sieve
- Powder Molecular Sieve 13x
- Molecular Sieve 3A
- Molecular Sieve 4A
- Molecular Sieve 5A
- Pharmaceutical desiccant
- Desiccant Canister
- Desiccant Packets
- Desiccant unit packs
- Desiccant Pillow packs
- Oxygen Absorbers
- Humidity Indicating Card
Access TOC: http://www.marketresearchfuture.com/ask_toc_request/us-europe-asia-pharmaceutical-desiccant-market-analysis-features-segmentation-global-market-forecast-to-2023
Regional Analysis
The packaging industry is gaining more importance globally, the reason behind this is the importance of protecting each and every product that is manufactured. The pharmaceutical desiccant industry is one of the emerging or booming industries globally.
The desiccant market for Europe is an emerging market and has experienced modest growth in recent years, the positive trend have continued but in 2015 there were a modest slowdown in the early half which was thought to have a decline in the market but in the send half of the year resulted to an average growth, thus the European market for desiccants is expected to be growing market.
The desiccant market for Asia is an market which stared in late, the market has huge potential as most of the emerging countries like India, China, etc are developing countries with developing markets, infrastructure and huge scope for FDI's thus this markets has huge potential for growth.
Talk to our Analyst : http://www.marketresearchfuture.com/schedule_call/us-europe-asia-pharmaceutical-desiccant-market-analysis-features-segmentation-global-market-forecast-to-2023
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- Spain
- Rest of Western Europe
Asia – Pacific
- Asia
- China
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- Japan
- South Korea
- Rest of Asia
- Pacific Countries
- Australia
- New Zealand
Middle East & Africa
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Poll: Do you think Claire le Gosse de Vampire should continue? Vote Now!
Author has written 1 story for Twilight.
Hello, my name is CloverofReason, though my real name is Imani. The name Imani means faith and it is actually the last day of Kwanzaa.
Right now I am working on a series called Claire le Gosse de Vampire which means "Claire the Vampire Brat". Obviously, I love languages and right now I am learning French and Japanese. Hopefully when I'm older I will be a Linguist or an Actress (the actress option is pretty random, I know).
I like to read, write, and draw but most of all I like to read! I can't draw for my life but my mom's an artist and I learned to love it. Writing is a personality but I never really thought I was very good at it. Claire le Gosse de Vampire is just a 'test drive' I don't know if it's really going to work out or not, but if you like it, THIS is why I like the reviews!

subset: commoncrawl


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_Mick_Phot... Photographer
Shure NekoleModel Female East Saint Louis, Illinois, US
My MM URL: http://www.modelmayhem.com/Shure1
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Recent Posts
Tag Archives: alarm clocks
True Story Tuesday: In which I finally get some sleep
Last May, I had a heart attack. Hint: I didn't die. But if I hadn't thought "this couldn't be a heart attack….could it?" and used the internet to figure out it probably WAS, and called 911, I wouldn't be here … Continue reading
Posted in Personal Tagged alarm clocks, Fatigue (medical), Health, heart, heart attack, Insomnia, Myocardial infarction, Sleep, worry Leave a comment


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I have a new client who has a proprietary program that was developed for him back in 1998. The program was written using Microsoft Access 97, and runs in the Access 97 Runtime mode. Due to various issues, the developer never put any maintenance functions into the program. Thus, all maintenance must be done via Access 97. The only computer my client had left with Access 97 on it just crashed and the original install CD are long since lost. Would anyone know where I might be able to download a copy of Access 97 at no cost? We have the key, so it would be legitimate install...we just need the install media.
Thank you in advance.
2 Replies
Jan 19, 2009 at 4:06 UTC
Short of recommending the various P2P networks, you might try to find either a pawn shop or a used software vendor in your area.
Someone like Bookman's Used Books Music & Software, 1056 S Country Club Dr
Mesa, AZ 85210, (480) 835-0505 or Computerland, 1245 E Northern Ave, Phoenix, AZ 85020 (602) 331-5030 may have what you seek.
Jan 19, 2009 at 4:19 UTC
Good options. I will research them and post back.
This discussion has been inactive for over a year.
You may get a better answer to your question by starting a new discussion.


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- Leading futurist Daniel Burrus, Chief Executive Officer of Burrus Research
- Marcus Buckingham, the best-selling author and former Gallup researcher who helped create the StrengthsFinder personal assessment tool used by millions
- Scott Hess, senior vice president, Human Intelligence at Spark Communications, a leading expert on the Millennial generation
- Youngme Moon, Senior Associate Dean and MBA Program Chair at Harvard Business School, and author of the best-selling book, Different
Fiserv, Inc. (NASDAQ: FISV), a leading global provider of financial services technology solutions, today announced the full agenda for Forum 2013 to be held April 15-18 at The Venetian and The Palazzo in Las Vegas. In addition to keynote speaker Gen. Colin Powell, who will share key leadership and life experiences, the newly announced speakers will provide insights on strategy, management and the trends that are impacting the next generation of financial services. The powerhouse lineup of speakers includes:


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Basket Set Round Blue Sapphire Stud Earrings
Round and Oval Blue Sapphire Dangle Earrings with Diamond Accents
3 Prong Martini Set Round Blue Sapphire Stud Earrings
Bar Set Round Diamond Halo Blue Sapphire Stud Earrings
Oval Sapphire Dangle Hoop Earrings in Open Diamond Frame
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4 Prong Set Blue Sapphire Basket Stud Earrings with Diamond
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Half Eternity Five Stone Blue Sapphire Wedding Band
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Prong Set Sapphire Bypass Ring with Diamond Accents
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Round Sapphire Solitaire Pendant with Diamond Accents
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6 Prong Tapered Shank Oval Solitaire Sapphire Ring
You have been with each other through thick and thin for fifty one years, and it's time to celebrate this proud moment. Give a precious blue sapphire wedding anniversary gift to your wife - a ring, pendant or a set of earrings. Browse Angara's blue sapphire jewelry collection that includes ornate, vintage, modern and subtle designs.
Bracelets studded with blue sapphires and diamond accents look spectacular and make unique presents on this significant occasion. At Angara, a variety of tennis bracelets and station bracelets are available. The sapphires are held amid shimmering halos and diamond stations enhancing the gemstone's intense blue allure. Sapphire rings are designed with alluring floral patterns, intricate scroll work on the gallery and elegant shank styles like a bypass, crossover or a split shank for a refined look. You can either go for simple and stackable sapphire rings or the exquisite and ornate ones.
If you are a son or daughter planning a grand anniversary for your parents, you can include a pair of gorgeous sapphire earrings as a 51st wedding anniversary gift for your mom. Know her taste for gemstone jewelry and shop the earrings that will perfectly match her personality. We have solitaire sapphire earrings designed with glimmering diamond halos, dainty drop earrings with lever backs and elaborate dangling earrings that make a luxurious style statement.
Sapphire pendants are also a great option for anniversary gifting. While the infinity knot and open circle pendants extend meaning to your gesture, pendants will floral halos, diamond encrusted bales and the stylish sapphire and diamond necklaces are pure indulgence. At Angara, we offer stunning sapphire anniversary gifts in a variety of stone qualities, metals, and total carat weight options so that you can easily choose the one that meets your style requirements and budget.


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Fatwire Article Portlet
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Arlington County is a county in the Commonwealth of Virginia. It is coterminous with the U.S. Census Bureau-census-designated place of Arlington, which is the second-largest principal city of the Washington metropolitan area. As a result, the county is often referred to in the region simply as "Arlington" or "Arlington, Virginia". In 2014, the county's population was estimated to be 229,302, which would make it the fourth-largest city in the Commonwealth if it were incorporated as such.
- Independent Professional Wrestling Alliance (1995-1999)
External links
- Arlington County, Virginia on Wikipedia


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HBE 5% NSR areas
at the Copper Mountain Mine
click here for more info
As of January 27th, 2015
Issued and Outstanding 58,159,989
Options Outstanding 4,300,000
Warrants Outstanding 0
September 2013
Hornby Bay Coppermine
property - 1.5mb
Western Nunavut
property overview- 8.5mb
All financial and news information concerning Hornby Bay Mineral Exploration can be found on SEDAR at the following link: Click Here For Sedar Information
Hornby Bay Mineral Exploration Ltd. (HBME) with its head office in Toronto, Ontario is a uranium exploration and development company with all of its uranium mineral properties located in Nunavut, Canada, which is the only Canadian jurisdiction that has settled its native land claim issues.
HBME was incorporated in 1996 under the Business Corporations Act (Ontario) as a uranium exploration company. From 1996 to 1998, HBME acquired mineral claims in the Hornby Bay Basin, which is located in western Nunavut and is one of the three major Proterozoic uranium basins in western Canada. Since 2003, the Company's overall strategy has been based on the systematic acquisition of relevant geophysical datasets and ground exploration activities for the definition of key target areas on its uranium properties. The uranium models pursued by HBME on its uranium properties include high-grade unconformity deposits; large, lower-grade disseminated deposits in sandstone; basement hosted high-grade, structurally controlled deposits; and vein type deposits.
The Company's overall strategy is based on:
The Company is a reporting issuer in Ontario, British Columbia and Alberta and trades on the TSX Venture Exchange under the symbol HBE.
More details regarding the company can be found by reviewing the documents currently on file with SEDAR [System for Electronic Document Analysis and Retrieval] www.sedar.com
Hornby Bay Mineral Exploration © 2010 | Powered by www.RMC.mobi


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Sorrel (Red Veined)
One of the most popular and visually striking leaves around. Red Vein Sorrel, also known as "hearts on fire", has a bold red stem and veins that run up the flatter but structured leaves. The flavor is sour and can be powerful depending on leaf size. Used sparingly it can provide a beautiful and flavor-balancing aspect to a plate. When out of stock, this product can be grown to order in 7 Weeks.
Sorrel growing instructions, cooking tips and recipes
Sorrel Red Veined | Salad Leaves
Red-veined dock Plant Care & Growing Information
Sorrel : Substitutes, Ingredients, Equivalents
How to Grow Sorrel
Beet-and-Red Sorrel Salad with Pistachio
Polish Sorrel Soup
Chickpea and Sorrel Salad
Creamy Salmon & Sorrel Recipe
White Peach and Sorrel Salad with Honey/Balsalmic Vinaigrette
More Info about Sorrel (Red Veined)
Sorrel (Red Veined) Recipes


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Soy blended candle with essential oils that will become a favorite.
No.02 Le Long Fond This scent is a savory mixture of hinoki wood accord underlined by notes of cedarwood and patchouli with a strong amber character.
Top Note: hinoki wood
Mid Note: cedarwood, patchouli
Base Note: white musk
No.04 Bois de Balincourt This scent is a sandalwood fragrance with a dominant cedarwood & sandalwood accord supplemented by a spicy cinnamon nutmeg complex with an earthy vetiver note
Top Note: sandalwood, cedarwood
Mid Note: vetiver, nutmeg, cinnamon
Base Note: amber wood
Top Note: Grapefruit, Orange, Black Pepper
Mid Note: cedarwood, Patchouli, Geranium
Base Note: Vetiver, Amber, Musk, Benzoin
No.10 Aboukir fragrance has a spicy eucalyptus top note with a floral amber heart composed of jasmine, lavender, carnation and a rich amber accord. The base notes of the fragrance incorporate a definitive woody accord of patchouli, cedarwood and vetiver with a sweet tonka bean accord.
Top Note: Lemon, Nutmeg, Eucalyptus
Mid Note: Jasmine, Carnation, Amber
Base Note: Patchouli, Cedarwood, Vetiver
Andtidris-Cassis is a fruity fragrance that begins with a black pepper note enhanced by bergamot & cassis. It then develops a strong green accord with a white rose base and dries down to a warm oakmoss, tonka and clean musk base.Top Note: Cassis, bergamot, black pepper
Andtidris-Jasmine is a romantic and gentle floral fragrance incorporates top notes of Italian orange and a rich green floral accord. The body of the fragrance is a floral bouquet of jasmine, ylang-ylang and tuberose. The fragrance dries down to an oakmoss and white musk background.Top Note: Italian orange


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IP Lookup
Simple current IP lookup, with geo-data.
The Unserializer
Convert serialized array or object data into easy-to-read formats.
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Current geomagnetic observations monitoring disturbances in the Earth's magnetic field and its affect on the GPS constellation.
Philips Hue
My dealings with Philips Hue smart home devices including a WordPress Plugin for interacting wit Philips hue Lights, Switches and Sensors.
Ding Ding
3DR Distributors
A list of 3D Robotics distributors/resellers including product range and service range (e.g. US/Canada, Europe/Schengen only, etc.).


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Missions of Space
Project Mercury began in 1958 and completed in 1963. Project Mercury was the first major U.S. Program.There goals were to successfully orbit earth with manned spacecraft,to investigate human kinds ability to function in space, and to recover both occupant and space craft. Each space craft carried one astronaut. There were seven Mercury astrounats, all military test pilots. Six of them flew aboard the Mercury space craft. The seventh was grounded due to minor medical aliment, but eventually flew as crewman aboard the Apollo-Soyuz Test project in 1975.
Project Gemini was the second human space flight program initiated by the United States. Operating between Mercury and Apollo, project Gemini consisted of 10 successful flights in 1965 and 1966. It provided crucial experience by developing and testing technics for advanced space travel and long-duration space operations necessary to land humans on the Moon. Gemini missions involved the first American extravehicular activity and new orbital maneuvers including rendezvous and docking. The objectives of the Gemini project were to gain astrounat experience with long duration test flights. Perfect methods of reentry and landing the spacecraft. Study the effects of weightlessness on astronauts during long flights.


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Question: When should you create a scenario instead of a data
When should you create a scenario instead of a data table? Give an example of a business situation that could best be analyzed with scenarios.
View Solution:
View Solution:
Answer to relevant QuestionsHow do pie charts differ from doughnut charts? What is a scenario in Solver? Describe the steps for saving a Solver model. What is the advantage of saving a Solver model? When should you include integer constraints in a Solver model? What is the disadvantage of using an integer constraint? Describe two methods to document information in a workbook.
Post your question


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The objective of milling gold ores is to extract the gold for the highest financial . For the grinding and crushing circuit at Goldstrike, Barrick relies on primary.Online Consultation


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Pazzo Gelato
3827 Sunset Blvd., Ste. D (Hyperion Ave.) Send to Phone
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OpenOpen daily
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* Click here for rating key
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Find out which restaurants and chefs top the list of GAYOT's 2016 Annual Restaurant Awards.


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Question: What is a virtual database What are the advan tages
What is a virtual database? What are the advan tages and disadvantages of using a virtual database?
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Castelinho's Homes
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About Castelinho's Homes
Truly, salvage from a daily schedule is required and that is where tourist destinations come into existence. Amongst a number of famous places, lies Goa, a land which is famed for its historic imprints revealed on every churches and temple present there. To adore this beauteous land and its facets, a visit to this place is must, and on the expedition one can find a sojourn at Castelinho's Homes. This property comes under the budget hotels in the vicinity offering a gamut of services and facilities. It is close to bus stand, railway station and other local transports. Apart from this, it is within reach to Anjuna beach and Mapusa. Facilities offered by the hotel include front desk, room service, backup generator and many more. Certainly, this budget hotel in Goa is worth a stay for all the visitors on a tight budget.
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See Photo Of Woman Who Neglected Her Baby At The Airport To Attend To Her Phone (Photo)
The picture of a mother has gone viral as she was seen at the airport neglecting her own baby, to pay more attention to her mobile phone.
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02/08/2017 Total Views: 10933 Today: 6
Verona. The property is 8.8 km from Sirmione and free private parking is offered. The apartment is composed of 1 bedroom and 1 bathroom, fitted with a bath or shower. A refrigerator and a stovetop can be found in the kitchen. Other facilities at Apartment Soiano del Lago 3 include a year-round outdoor pool. Riva del Garda is 47.6 km from Apartment Soiano del Lago 3, while Mantova is 46.5 km away. The nearest airport is Villafranca Airport, 33 km from Apartment Soiano del Lago 3. (Apartment 3 Soiano del Lago hotels booking).


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England's top-flight will soon reconvene for the 2014-15 season, with Manchester City the defending champions and Leicester City, Burnley and QPR the promoted teams
As the campaign gets underway, Louis van Gaal's Manchester United will face Swansea, Mauricio Pochettino's Tottenham take on London rivals West Ham and Arsenal take on Crystal Palace.
Elsewhere, Leicester face Everton, QPR take on Hull, Stoke clash with Aston Villa and West Brom take on Sunderland.
The following day, two more games will be played, with Liverpool hosting Southampton and Manchester City travelling to Newcastle, before Chelsea's clash with Burnley rounds off the weekend's action.
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Mango and Pineapple Jam
Recipe by Sharon123
This is yummy and makes a tasty topping for vanilla ice cream.
Top Review by Abby Girl
This is a fast and easy recipe to put together and I ended up with 4 half pint jars. As it was, I had some ripe mango and some lime to use up and just happened to have a can of pineapple in the cupboard. I took the other reviewer's suggestions and cut the sugar down to 1 cup. I may have overcooked the jam a bit because it came out very, very thick. Do try it over ice cream...it is quite the treat. I mix some of the jam with some brandy to thin it out and heated it up a bit over the stove...then poured over the ice cream. I think giving it a water bath might be a good idea as I had 2 jars that did not seal. I would do this recipe again!!!
See Discussion:22 Reviews
Ingredients Nutrition
- 2 cups ripe mangoes, cut up
- 1 cup crushed pineapple
- 1⁄3 cup lime juice (or lemon)
- 2 cups sugar
Directions See How It's Made
- Peel mango and cut in small pieces, then mash with potato masher to bring juice out.
- Cook 5 minutes.
- Add rest of ingredients, mix well and bring to a boil.
- Boil rapidly 15 to 20 minutes, stirring often, until thick.
- Pour into small sterilized jars.
- Seal and store in cool place.


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Gone are the days when our uncles used to pull up a deck of cards to show us our favourite magic tricks. But for most of us, the fascination for magic tricks isn't. We all want to learn how themystery magic tricks were performed, right? After all, who does not want to be the coolest guy or girl in his or her school and college? For More :- http://www.magicmysteryapp.com/\n
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In its first year the Red & Blue Rewards program had over 400 members who racked up 3,450 points for the entire year. Students gained points by attending football, basketball, wrestling, and select non-ticketed sports.
The rewards program isan initiative forPenn students only, with the goal of getting more students out to Penn Athletics events over the year. Members won prizes for reaching different point totals and the highest point winner for the year was rewarded a free IPod Touch. The winner of the IPod touch was Matthew Sommer with a total of 66 points for the year.
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Stabenow: Senate-Approved Version of Transportation Bill Provides Billions More for Michigan Roads, Creates Michigan Jobs
The state of Michigan, hard hit by job losses, could see 54,000 good-paying jobs coming its way with the Senate's passage of a multi-year transportation bill, U.S. Senator Debbie Stabenow (D-MI) said today.
The bill, provides $295 billion in highway and transit funding - $11 billion more than the companion House version. "This highway bill is good for Michigan - good for Michigan roads, Michigan transit systems and Michigan jobs," Stabenow said. "This bill has passed the Senate with strong bipartisan support, and now I will work to protect the important gains we have made for Michigan in the Senate version of this bill, as this legislation moves to conference with the House."
Stabenow said the estimate of 54,000 new jobs in Michigan is based on a U.S. Department of Transportation estimate that each $1 billion in spending creates 47,500 new jobs and generates more than $2 billion in economic activity. Michigan is estimated to receive an average of $1.113 billion in spending each year over the next five years, for a total of $5.66 billion.
"Over the last four years Michigan has lost jobs, and this transportation bill will create jobs - good-paying jobs that will help thousands of Michigan families pay their mortgages, save for retirement, and pay for the children's college tuition costs," Stabenow said.
Stabenow played a key role in giving final shape to the Senate version of this transportation bill, working with Sen. Jim Talent (R-MO) on a bipartisan amendment that was included in the Senate's final budget resolution and helped pave the way for the additional $11 billion added to the Senate bill.
The Senate version of the SAFE-TEA transportation bill improves funding for Michigan by $51 million per year above what the U.S. House approved, but it even more dramatically improves the state's position over the prior multi-year transportation bill, TEA-21, increasing annual highway and bridge funding by $249 million per year.
Under the Senate version of the bill, Michigan can expect to receive more than $600 million in total transit investments, an increase of $173 million over TEA-21.


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The point of sale is the pressure point of the supermarket. At the same time as retailers load it up with new customer payment options, marketing messages and data-gathering devices, they also want accurate, quick customer throughput -- all using technology that's simple enough for even a minimally experienced cashier to operate.
It's no wonder that retailers are seeking ways to combine and simplify POS functions, by more closely integrating payment solutions and using wireless techno


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Regular "mucosal block" is characterized by decreased uptake of a normal iron load 3–72 h after the administration of excess iron (generally 10 mg) to iron-deficient animals. We found that short-acting mucosal block could be induced by much lower iron concentration and much shorter induction time than previously reported, without affecting levels of gene expression. A rapid endocytic mechanism was reported to decrease intestinal iron absorption after a high iron load, but the activating iron load and the time to decreased absorption were undetermined. We assessed the effects of 30–2,000 μg iron load on iron uptake in the duodenal loop of iron-deficient and iron-sufficient rats under anesthesia. One hour later, mucosal cellular iron uptake in iron-deficient rats administered 30 μg iron was 76.1%, decreasing 25% to 50.7% in rats administered 2,000 μg iron. In contrast, iron uptake by iron-sufficient rats was 63% (range 60.3–65.5%) regardless of iron load. Duodenal mucosal iron concentration was significantly lower in iron-deficient than in iron-sufficient rats. Iron levels in portal blood were consistently higher in iron-deficient rats regardless of iron load, in contrast to the decreased iron uptake on the luminal side. Iron loading blocked mucosal uptake of marginally excess iron (1,000 μg), with a greater effect at 15 min than at 30 min. The rapid induction of short-acting mucosal block only in iron-deficient rats suggests DMT1 internalization.
- iron absorption
- mucosal block
- iron deficiency
- duodenum
- excessive iron loading
- ferritin
iron deficiency induces the upregulation of several intestinal iron transporters, including divalent metal transporter 1 (DMT1, formally Slc11a2) and ferroportin (5, 6, 10, 13, 17, 29, 32), which increase iron absorption in the small intestine. However, excess iron loading suppresses intestinal iron absorption, a phenomenon known as "mucosal block" and found to persist for several hours to several days (4, 11, 23, 25). The effect of oral ingestion of 10 mg iron on the duodenal expression of molecules associated with intestinal iron transport was tested in iron-deficient rats and related to the mucosal block phenomenon (8). Three hours after oral iron ingestion, 59Fe uptake (the counts in the carcass and duodenum) and transfer (the counts in the carcass) decreased 60 and 50%, respectively. This inhibition of iron absorption is accompanied by the downregulation of DMT1, duodenal cytochrome b (Dcytb, formally Cybrd1), and ferroportin 1 (FPN1, formally Slc40a1) mRNA and protein (8). Generally, oral administration of 10 mg iron is considered excessive in experiments examining intestinal iron absorption. We found that direct application of 10 mg iron to the loop of the duodenum of iron-deficient rats downregulated Dmt1 mRNA and upregulated ferritin (Ft) mRNA 2 h later, with a longer time (>2 h) required for FT protein synthesis in the duodenal mucosa (1).
Because the mucosal block phenomenon is usually examined in iron-deficient animals, increased intracellular iron concentration in the small intestines following iron loading may be due to the increased expression of DMT1 protein. Regardless of whether mucosal block occurs only in iron-deficient rats, the iron concentration required to induce this phenomenon and the time to onset remain unclear. We therefore administered 10 mg of iron to the ligated duodenum of iron-deficient and iron-sufficient rats and measured their serum iron and duodenal iron uptake. One hour later, arterial serum iron concentration was higher in iron-deficient (928.9 ± 100.3 μg/dl) than in iron-sufficient rats (642.2 ± 30.6 μg), suggesting increased iron absorption in the former, but duodenal iron uptake did not differ significantly (5,658 and 7,533 μg/h, respectively). On the basis of these results, we hypothesized that short-acting mucosal block occurs prior to the regular mucosal block, which is characterized by the downregulation of mRNA and proteins. To determine whether an activating iron load can induce a short-acting mucosal block, we assessed iron uptake from the lumen of the small intestine following duodenal administration of iron loads of 30 μg (low) to 2,000 μg (marginally excessive).
DMT1 was also internalized in the duodenum of these rats several hours after oral loading of a very high concentration of iron (8). Administration of an excess iron load to the stomach of iron-deficient rats reduced duodenal DMT1 protein expression and apical distribution, as well as reducing fluorescence intensity in the cytoplasm (20). Furthermore, DMT1 was internalized by Caco-2 cells within min (20). We therefore assessed whether short acting mucosal block occurs only in iron-deficient rats and also analyzed the time to onset in iron-deficient duodenal mucosa, by comparing the effects of administering FeSO4 to the duodenal loop of iron-deficient and iron-sufficient rats before the decreases in gene and protein expression (i.e., <2 h).
Experimental animals and diets.
Weaned male Wistar rats (4 wk old) were obtained from CLEA Japan (Tokyo, Japan) and fed an AIN-93G iron-sufficient diet (24) or an iron-deficient diet for 3 wk. The AIN-93G diet (CLEA Japan) contained 35 mg/kg iron as FeC6H5O7·nH2O, whereas the iron-deficient diet lacked FeC6H5O7·nH2O. Experimental diets and deionized water were provided ad libitum. The iron contents of the iron-sufficient diet, the iron-deficient diet, and deionized water were 54 mg/kg, 5 mg/kg, and <1 ppm, respectively, as determined by atomic absorption spectrophotometry (AA-7000; Shimadzu, Kyoto, Japan). The rats were housed individually in stainless steel cages maintained at a constant temperature (23 ± 2°C) and humidity (50 ± 10%) with a 12-h light-dark cycle.
Rats were anesthetized by intraperitoneal injection of pentobarbital (0.1 ml/100 mg body wt; Dainippon Pharma, Osaka, Japan), and a midline incision was made in the abdomen. The duodenum was exposed, and two ligatures were tied with disposable microvascular clips (BEAR Medic, Ibaraki, Japan), one ∼0–1 mm from the pylorus and the other at the ligature of Treitz, ∼6–8 cm from the pylorus. Incisions were made proximal to the upper ligature and distal to the lower ligature. The small intestine between the upper and lower ligatures was flushed with prewarmed saline (37°C) from the upper incision. The microclips were moved to seal the pylorus and the distal ends of the duodenum, and the loop was instilled with 10 mM HCl (control) or iron test solution. In experiment 1, the test solution contained 30–2,000 μg iron as FeSO4 (Wako Pure Chemical, Osaka, Japan) in 1 ml of 10 mM HCl. The control and iron solutions contained 0.5% polyethylene glycol (PEG; ICN Biomedicals, Aurora, OH) as a soluble absorption marker. The abdominal incision was sutured with thread and covered with gauze soaked in prewarmed saline (37°C) to prevent the wound from drying. The rats were kept warm for the indicated times to avoid hypothermia. One hour after infusing the test solution, blood was collected from the portal vein and the descending aorta and centrifuged, and serum was separated to measure iron concentration and unsaturated iron binding capacity. The duodenum was then carefully removed from each rat. The duodenal contents were collected, the lumen was rinsed with saline, and the mucosa was scraped off. Mucosal samples were immediately frozen on dry ice and stored at −80°C until use. To estimate iron retention in the duodenal mucosa 1 h after loading, the duodenal loops were collected and washed with 10 mM HCl to remove iron from the mucosal surface.
In experiment 2, the iron solution contained 100 or 1,000 μg of iron as FeSO4. Blood from the portal vein and the descending aorta, as well as the duodenal contents and mucosa, were collected 15 or 30 min after loading. To estimate iron retention and ferritin protein expression in the liver 15 and 30 min after loading, the livers were collected and blood in the liver was replaced with saline. The other experimental procedures were the same as those in experiment 1, unless otherwise noted. All blood/tissue sampling was done between 05:00 and 12:00, when the rat serum iron concentrations were low, to avoid the effects of diurnal variation (2) on serum iron. To prevent iron contamination from the environment, all instruments used to prepare the test solutions and mucosal samples were soaked in 1 M HCl and washed with ultrapure water. The Ethics Committee of Tokyo Metropolitan University approved all animal procedures.
Determination of iron uptake and iron concentrations in the duodenum and the liver.
The concentrations of iron in the test solutions and in the duodenal contents were immediately quantified by atomic absorption spectrophotometry. Precipitates were dissolved by dropwise addition of 1 M HCl. The amount of PEG was estimated by the nephelometric method at 540 nm after deproteinization of the samples (14). Iron uptake was determined by using the formula: Iron uptake (μg) = Fe in iron test solution (μg/5 mg PEG) − Fe in the duodenal content (μg/5 mg PEG). The percentage uptake was calculated as the ratio of iron uptake to iron load. To estimate iron retention in the duodenal mucosa and the liver, the duodenal loops and the liver homogenates were air dried and dried to ash at 450°C. Iron content was then determined by atomic absorption spectrophotometry.
Determination of ferritin protein in the duodenum and liver.
The duodenum and liver were homogenized in bioMasher (Nippi Incorporated Protein Engineering Office, Tokyo, Japan) and FT protein was extracted with RIPA buffer (Thermo Scientific, Vernon Hills, IL). FT proteins were quantified by Rat Ferritin ELISA (Immunology Consultants Laboratory, Portland, OR). Total proteins in duodenum homogenate were quantified by BCA protein assay.
Serum iron concentration, TIBC, and transferrin saturation in portal and arterial blood.
Serum iron concentration and unsaturated iron binding capacity (UIBC) were measured on a 7170 Clinical Analyzer (Hitachi High-Technologies, Tokyo, Japan), by the nitroso-(N-propyl-N-sulfo-propylamino)phenol method. Total iron binding capacity (TIBC; μg/dl) was calculated as the sum of the serum iron concentration and UIBC (the amount of iron able to bind unsaturated part of transferrin.). Transferrin saturation was calculated as serum iron concentration divided by TIBC concentration.
Expression of Dmt1, Ft, and Fpn1 mRNAs.
Mucosal samples were homogenized and total RNA was extracted from 10 mg of each using a QuickGene RNA tissue kit S (RT-S2) and the QuickGene-Mini 800 (Fujifilm, Tokyo, Japan). DNase solution (Nippon Gene, Tokyo, Japan) was added to each sample, and the samples were incubated at room temperature for 5 min. Each total RNA sample had an A260/A280 between 1.7 and 2.0. RNA integrity was assessed with an Agilent 6000 RNA Nano Kit and a Agilent 2100 Bioanalyzer with 2100 expert software. cDNA was synthesized from 5 μg of each total RNA by using a PrimeScript RT reagent Kit according to the manufacturer's instructions (Takara Bio, Shiga, Japan). The relative levels of expression of Dmt1, Ft (heavy chain), and Fpn1 mRNA were determined by real-time PCR (Chromo4 Real-Time PCR Detection System; Bio-Rad Laboratories, Hercules, CA) with TaqMan probes and iQ Supermix (Bio-Rad Laboratories). Relative expression was determined by using calibration curves, with stock cDNA from the duodenal mucosa of iron-deficient rats. Transcript levels were normalized relative to the expression of glyceraldehyde 3-phosphate dehydrogenase (GAPDH) mRNA in that sample, and divided by the mean level in the control group. Primer sequences and probes are shown in Table 1, and the PCR methods, except for Fpn1, have been described (1). Fpn1-specific primers and probe were designed by use of Universal ProbeLibrary Assay Design Center software (Roche Diagnostics, Basel, Switzerland). The amplification protocol for Fpn1 consisted of an initial denaturation at 95°C for 10 min, followed by 40 cycles of denaturation at 90°C for 10 s, annealing at 55°C for 1 min, and extension at 72°C for 30 s. All reactions were performed in triplicate and the data are presented as means ± SD.
Statistical analysis.
All results are reported as means ± SD for 4–7 rats. Student's t-tests were used to assess differences between rats fed the iron-deficient and iron-sufficient diets at the same iron load in experiment 1. Significant differences among the iron loads in rats fed the iron-deficient and iron-sufficient diets were determined by one-way or two-way ANOVA followed by Tukey's post hoc test. Differences in gene expression between the control and experimental groups were determined by Dunnett's multiple comparison tests. Differences were considered significant at a P value < 0.05.
After 3 wk of feeding, mean ± SD body weights and feed efficiency were significantly lower in rats fed the iron-deficient than the iron-sufficient diet, both in experiments 1 and 2, due to decreased feed intake by iron-deficient rats (Table 2).
Serum iron concentrations and transferrin saturation of portal and arterial blood (experiment 1).
Table 3 shows the serum iron concentrations, UIBC, TIBC, and transferrin saturation of portal blood in iron-deficient and -sufficient rats. Serum iron concentrations were significantly lower in iron-deficient than in iron-sufficient rats administered vehicle (control). Iron saturation markedly decreased as TIBC increased, a phenomenon usually associated with increased iron absorption. We therefore administered various concentrations of nonheme iron (i.e., FeSO4) to the duodenal loops of rats and measured serum iron concentrations in portal blood 1 h later. Iron loading (30–2,000 μg) into the duodenal loops proportionately increased serum iron concentration, with transferrin saturation reaching nearly 100% at iron loads ≥1,000 μg, with no large variations in TIBC. Serum iron concentrations were consistently higher in iron-sufficient than in iron-deficient animals at iron loads ≤30 μg but were lower in iron-sufficient rats at iron loads ≥500 μg. The differences in serum iron contents between rats administered vehicle and 30 μg iron were 26 μg/dl (244.5-218.1 μg/dl) in the iron-sufficient group and 94 μg/dl (170.3-76.4 μg/dl) in the iron-deficient group, indicating that iron absorption was greater in the iron-deficient rats. At iron loads ≥200 μg, the serum iron contents increased 210–302 μg in iron-sufficient and 418–748 μg in iron-deficient rats. At iron loads of 200, 500, 1,000, and 2,000 μg, the increases in iron contents in the iron-deficient group were 418 μg (494.7−76.4 μg/dl), 560 μg (636.5−76.4 μg/dl), 713 μg (789.6−76.4 μg/dl), and 748 μg (824.8−76.4 μg/dl), respectively. Therefore, greater iron loads were associated with greater differences in serum iron contents between iron-sufficient and iron-deficient rats.
Experimental results of arterial blood are shown in Table 4. Hematocrits were significantly lower in iron-deficient (23.5–27.8%) than in iron-sufficient rats (43.9–49.3%). The effects of iron loading on serum iron concentrations, TIBC, and transferrin saturation in arterial blood were consistent with those of portal blood, despite passing through the liver.
Duodenal iron uptake 1 h after luminal administration of a single iron dose (experiment 1).
Iron uptake was higher in iron-deficient than in iron-sufficient rats after loading with 30 and 200 μg iron but did not differ after loading with 500 μg iron (Table 5). Iron uptake after loading with 1,000 and 2,000 μg was significantly lower in iron-deficient rats than in iron-sufficient rats. At 1 h, the percent iron uptake in iron-sufficient rats was 63% (range 60.3–65.5%), regardless of iron load. The percent iron uptake in the iron-deficient group was 76.1% at 30 μg, decreasing approximately 25% to 50.7% at 2,000 μg.
Iron accumulation in the duodenum.
Iron accumulation in the duodenum 1 h after iron loading is shown in Fig. 1. After loading with vehicle, the mean duodenal iron concentrations in the iron-sufficient and iron-deficient groups were 14.4 ± 1.3 and 6.9 ± 0.6 μg/100 mg dry tissue, respectively. Iron accumulation tended to be lower in the iron-deficient than in the iron-sufficient group at any iron load, particularly at loads ≥500 μg.
Serum iron concentration and transferrin saturation in portal and arterial blood (experiment 2).
The duodenal loops of iron-deficient and iron-sufficient rats were administered 100 μg or 1,000 μg of nonheme iron, and portal and arterial blood were obtained 15 or 30 min after loading. Tables 6 and 7 show the serum iron concentration, UIBC, TIBC, and transferrin saturation of portal and arterial blood, respectively. After an iron load of 100 or 1,000 μg, the portal and arterial serum iron concentrations were higher in the iron-sufficient than in the iron-deficient group, except for 30 min after an iron load of 1,000 μg. However, the increments in serum iron concentrations were higher in the iron-deficient group at all iron loads. The increases in serum iron concentration were estimated as differences, by subtracting serum iron concentrations in vehicle-loaded from iron-loaded animals. For example, at an iron load of 100 μg, the incremental iron concentrations at 15 min were 320.9-225.5 μg/dl (control) = 95.4 μg/dl in iron-sufficient and 179.2-23.2 μg/dl = 156.0 μg/dl in iron-deficient rats. At an iron load of 1,000 μg, the incremental iron concentrations at 30 min were 196.6 μg/dl (422.1-225.5 μg/dl) in iron-sufficient and 411.0 μg/dl (434.2-23.2 μg/dl) in iron-deficient animals. Table 7 shows that hematocrits were significantly lower in iron-deficient (20.4–23.1%) than in iron-sufficient (41.3–43.5%) rats. As observed in experiment 1, the effects of iron loading observed in arterial blood were consistent with the effects on portal blood.
Duodenal iron uptake after luminal administration of normal and marginally excessive iron loads (experiment 2).
Iron uptake was assessed 15 or 30 min after loading with 100 or 1,000 μg iron (Fig. 2). Fifteen minutes after a load of 100 μg iron, iron uptake was significantly higher in iron-deficient than in iron-sufficient rats. The percent iron uptake in the iron-deficient group tended to be higher at 15 min than at 30 min but was similar at both times in the iron-sufficient group. After a 1,000 μg load, iron uptake was significantly lower in the iron-deficient than in the iron-sufficient group, particularly at 15 min after loading.
Dmt1, Ft, and Fpn1 mRNA expression.
Following loading with vehicle, the level of Dmt1 mRNA expression in the duodenum was 3.9-fold higher in iron-deficient than in iron-sufficient rats (Fig. 3AI, P < 0.01), whereas the relative levels of ferritin (Fig. 3B) and Fpn1 (Fig. 3C) mRNAs in iron-deficient rats were 0.66 and 0.92, respectively, compared with iron-sufficient rats (not significant). Under these conditions, Dmt1 with iron-responsive element (IRE) and non-IRE were both upregulated in iron-deficient rats, with a greater change in Dmt1 with IRE (Fig. 3AII). As expected, a luminal iron load of 100 μg or 1,000 μg did not significantly alter the levels of expression of any of these genes in either group, relative to vehicle-treated rats.
Iron and ferritin contents in the duodenum and liver.
After loading with vehicle, the mean duodenal iron concentrations in the iron-sufficient and iron-deficient groups were 15.0 ± 2.6 and 7.0 ± 1.2 μg/100 mg dry tissue, respectively, consistent with the results of experiment 1 (Fig. 4). Following an iron load of 1,000 μg, iron accumulation tended to be lower in the iron-deficient than in the iron-sufficient group, although the difference was not significant. Duodenal ferritin, hepatic iron, and ferritin contents were significantly higher in iron-sufficient than in iron-deficient rats.
How does the short-acting mucosal block differ from the regular mucosal block?
Mucosal block is a phenomenon in which iron absorption is suppressed for several days after a large iron load, which was first confirmed in dogs in 1943 (11). Duodenal uptake and transfer of 59Fe after an oral load of 10 mg of nonradioactive iron was found reduced at 3 h and persisted for 12 h (8). Furthermore, DMT1 protein expression was decreased and FT protein increased, suggesting that mucosal block is characterized by suppressed iron uptake coupled with abnormal DMT1 gene and protein expression. This phenomenon has been attributed to excess intracellular iron levels caused by elevated iron uptake capacity resulting from the upregulation of DMT1 expression in the iron-deficient state. Hepatic hepcidin synthesis was enhanced by increased iron storage in the liver, causing FPN degradation on the basolateral surface of epithelial cells (16, 21, 28, 31).
The results presented here showed that the short-acting mucosal block, with no downregulation of mRNA, occurred prior to the induction of the regular mucosal block. The short-acting mucosal block could be induced by much a lower amount of iron (1,000 μg) and much faster (15 min) than previously reported. This transient suppression of iron uptake in response to high iron load was specific to iron-deficient rats, with no inhibition of iron absorption into the portal blood observed. Iron absorption must be enhanced in iron-deficient rats, because the increments in portal and arterial serum iron concentrations were higher in the iron-deficient group at all iron loads. However, mucosal iron accumulation tended to be lower in the iron-deficient than in the iron-sufficient group, particularly at excessive iron load, as a result of transient suppression of iron uptake and increased iron transport to portal blood.
Activating iron load and time to induce the short-acting mucosal block.
In iron-deficient rats, 70% of iron was taken up 1 h after loading with low iron loads, but decreased by 25% to 50% at higher, excessive loads, while being approximately 63% in iron-sufficient rats, regardless of iron load. The iron content in the duodenum was also lower in iron-deficient than in iron-sufficient rats 1 h after loading. Overall 59Fe absorption by jejunal segments from iron-deficient rats was found to be 2.0- to 4.6-fold higher in segments from iron-sufficient rats at luminal concentrations ranging from 1 to 100 μmol/l, much lower than the 0.5–35 mmol/l concentrations in our study (18). We found that duodenal iron uptake was suppressed at iron loads exceeding 500 μg. Because iron uptake was higher in iron-deficient rats than in iron-sufficient rats at iron loads <200 μg, the luminal iron concentration must be monitored in the duodenum, with suppression of iron uptake when iron concentrations exceed a threshold level. In ligated duodenum, the suppression of iron uptake was detected at iron concentrations between 200 and 500 μg/ml. Because the daily intake of iron by rats used in these experiments was approximately 500 μg, not only medical iron preparations but also dietary iron supplements may suppress iron uptake. Generally, a supplemental daily intake of one- to twofold amounts of dietary iron at one time is regarded as not harmful to the small intestine, but lower iron loads may be a burden on the small intestinal mucosa.
We also found that the reduction in iron uptake after loading with 1,000 μg iron was stronger at 15 min than at 30 min, suggesting that this suppressive response occurred very quickly, before changes in gene and protein expression. This is an important difference between the regular and the short-acting mucosal block. As we previously reported (1), significant decreases in Dmt1 mRNA expression were first observed 2 h after the administration of an excess iron load. As the blood vessels and biliary tract are preserved in this experimental model, the pH of the duodenal contents rose from pH 1 to pH 6–7 immediately after administration of iron in 10 mM HCl to the ligated duodenum. Therefore, iron uptake and iron transfer in our model reflect the actual environment of the small intestine.
Although the mechanism responsible for this suppression is not clear, it is important that suppression by marginally excessive iron was induced only in iron-deficient, not in iron-sufficient, rats. The short-acting mucosal block may protect mucosal cells from oxidative stress caused by excess iron, because FT, which chelates iron, has antioxidative properties, and its level of expression was extremely low in enterocytes of iron-deficient rats.
Levels of expression of genes and proteins are not changed by iron loading in the short-acting mucosal block.
The degradation of duodenal DMT1 protein was reported to require >1 h after oral administration of 10 mg iron (8). The findings reported here are unlikely to be due to the degradation of DMT1 protein because we observed suppression of iron uptake as early as 15 min after loading with 1,000 μg iron. Although iron uptake was strongly suppressed within 15 min after loading, the level of suppression declined over time, resulting in comparable rates of iron uptake by iron-sufficient and iron-deficient rats 30 min after loading. These findings suggest that the short-acting mucosal block involves a suppressive response mediated by DMT1 protein (e.g., internalization) that persists until the regular mucosal block (i.e., mucosal block accompanied by decreases in DMT1 gene and protein expression). The IRE in Dmt1 mRNA (5, 7, 27) may be involved in this short-acting suppression, because the level of Dmt1-IRE mRNA is upregulated 18.5-fold in the duodenum of iron-deficient rats. The IRE in Dmt1 mRNA has been reported induced by dietary iron deprivation (5). DMT1 proteins with distinct COOH-termini (15) have been reported translated from distinct mRNAs by alternative splicing of 3′ exons encoding different 3′ untranslated regions. During the transferrin cycle, DMT1 not containing IRE is recycled with transferrin receptor between the plasma membrane and endosome, whereas DMT1 containing IRE is not (26). However, this mechanism has not been confirmed in duodenal enterocytes. Iron has been reported to induce apical-basal movement of DMT1 in Caco-2 cell monolayers (20) and in rat intestines (20, 22, 30). Cellular internalization of DMT1 in the duodenum of Belgrade and Sprague-Dawley rats was observed 1–2.5 h after dietary iron intake and within 30 min in Caco-2 cells. As predicted by the membrane distribution of DMT1, high extracellular iron levels caused a robust decrease in apical iron uptake, suggesting that mucosal block may occur earlier in iron-deficient rats than previously thought. Therefore, the suppression of iron uptake 15–30 min after loading is likely due to a change in function of DMT1 protein, possibly due to the internalization of DMT1 in an apical-basal manner (19, 20). Possible differences in DMT1 localization, especially focused on DMT1 with and without IRE, should be assessed immunohistochemically, to determine whether DMT1 internalization is accompanied by transient suppression of iron uptake in vivo. The transport of iron to the portal blood after iron load was consistently higher in iron-deficient than in iron-sufficient rats, regardless of the iron load. Transient suppression of iron uptake was observed only at the apical surface (i.e., transport from the lumen into the mucosal cells), not at the basolateral surface (i.e., from the mucosal cells to the portal blood). Serum hepcidin levels in rats were observed to increase within 1 h after loading excess iron through a nasogastric tube, resulting in acute intoxication levels of 20 or 40 mg/100 g body wt, and were found to remain elevated for an additional 6 h (3). Therefore, the transient mucosal block would not involve FPN1 degradation by upregulated serum hepcidin.
Iron regulation of DMT1 has been reported to involve the tissue-specific expression of an upstream 5′ exon (exon 1A), with higher expression in the duodenum and kidneys than in other tissues (12). This exon 1A contains a conserved sequence of 29–31 amino acids, which may be required to induce the transient mucosal block. The cytoplasmic tail domain of IRE-less DMT1 was also found to determine the isoform-specific localization of DMT1 (26). Clearly, additional studies are needed to assess the roles of these sequences in the function of duodenal DMT1.
The luminal iron concentration must be monitored in the duodenum if iron concentrations exceed a threshold level, to induce the short-acting mucosal block. Thus our finding that the short-acting mucosal block was induced only in iron-deficient, not in iron-sufficient, rats was likely important. Dmt1-IRE mRNA is induced by iron deficiency, with the level increased 18.5-fold in iron-deficient duodenum. Moreover, DMT1 proteins with distinct COOH-termini containing IRE (15) have been reported. The iron-regulatory protein (IRP) is also expected to play key role in the mucosal block, since IRP deficiency impairs iron absorption and promotes mucosal iron retention via a ferritin-mediated mucosal block (9). Thus experiments are currently underway to quantitatively determine whether DMT1-IRE protein undergoes in internalization or functional changes in vivo.
This work was supported by a Grant-in-Aid (no. 22650182) from the Ministry of Education, Culture, Sports, Science and Technology of Japan (MEXT).
No conflicts of interest, financial or otherwise, are declared by the author(s).
S.S., S.Y., E.N., and A.A. conception and design of research; S.S., S.Y., E.N., K.T., and A.A. performed experiments; S.S., S.Y., E.N., and A.A. analyzed data; S.S. interpreted results of experiments; S.S. prepared figures; S.S. and A.A. drafted manuscript; S.S. edited and revised manuscript; S.S. approved final version of manuscript.
Present address for E. Nozaki: Department of Laboratory Medicine, Kyorin University, 6-20-2 Shinkawa Mitaka, Tokyo 181-8611, Japan.
Present address for K. Tadai: Department of Nutrition, Shubun University, 6 Nikko-cho, Ichinomiya, Nagoya 491-0938, Japan.
- Copyright © 2014 the American Physiological Society


text:
Spar Mid Wales Lge Two
NEWBRIDGE boosted their promotion bid with a 2-1 home victory over Abermule in Spar Mid Wales League Two.
Ricky Davies' strike was not enough for the visitors as Lyndon Jones and a Ryan Lawrence bullet header earned the Bridgemen three points and closed the gap on the pace setters to two points.
Meanwhile Henry Nicholson received top marks with a double as Aberystwyth University overcame Penybont 3-2 on Sunday.
Jamie Stuart completed the scoring for the Students while the hosts remained in contention throughout thanks to goals from Matthew Boore and Jake Bufton.
See full story in the County Times


text:
Look at other dictionaries:
swimmeret — 1840, from agent noun from SWIM (Cf. swim) (v.) + dim. suffix. Related: Swimmerets … Etymology dictionary
swimmeret — [swim′ər et΄] n. any of the small, abdominal appendages, or pleopods, in certain crustaceans, used primarily in swimming and for carrying eggs … English World dictionary
swimmeret — noun Date: 1840 one of a series of small unspecialized appendages under the abdomen of many crustaceans that are best developed in some decapods (as a lobster) and usually function in locomotion or reproduction … New Collegiate Dictionary
swimmeret — n. [A.S. swimman, to swim] (ARTHROPODA: Crustacea) An abdominal appendage functioning as a swimming organ; pleopod … Dictionary of invertebrate zoology
swimmeret — /swim euh ret /, n. (in many crustaceans) one of a number of abdominal limbs or appendages, usually adapted for swimming and for carrying eggs, as distinguished from other limbs adapted for walking or seizing. [1830 40; SWIMMER + ET] * * * … Universalium
swimmeret — noun In decapods such as lobsters, swimmerets, technically called pleopods, are legs primarily used for swimming but also used for brooding the eggs (except in prawns) and catching food … Wiktionary
swimmeret — See: pleopod [Martin, 2005]. See: pleopod [Butler, T.H.]. See^ pleopod [McLaughlin, 1980]. (Order Decapoda): Pleopod [Stachowitsch, 1992] … Crustacea glossary
swimmeret — swim·mer·et || swɪmÉ™ ret / swɪmÉ™ret n. any of the pair of small abdominal limbs or appendages in several crustaceans … English contemporary dictionary
swimmeret — [ swɪmərɛt] noun Zoology a forked swimming limb attached to a crustacean s abdomen … English new terms dictionary
swimmeret — swim·mer·et … English syllables


text:
login with
not too sure what to all put in my hospital bag?
I brought an outfit for me and baby to go home in, extra pair of undies and toiletries even though the hospital provided a lot of stuff for me there, thy also gave me a blow up doughnut to sit on for the ride home and ice pack pads for my privates. Make sure your car seat is ready too.
I read through so many lists. I brought SO much stuff with me and didn't end up using any of it. Let me tell ya hon, you need an outfit to go home in, something comfy. An outfit for baby to go home in, whatever works with the weather. A bag of toiletries. I brought a housecoat to walk around in (was in the hospital for 5 days so I got bored). Your camera, phone charger. The hospital will provide you with nearly everything so don't worry about diapers or wipes or anything for baby. If you're unsure call your maternity ward and they'll let you know what they will and won't give you. Also.. lol I know this sounds bad but it was advice given to me. The hospital is going to charge sooo much for his. So take everything. lol Take the wipes, take the diapers, take the little sample bottles of formula. And don't feel bad about it! haha.. good luck!
Moms Expertise10/14/14
Of my four kids, I think I opened my hospital bag once after I got there... that was to shove all the supplies (diapers, wipes) from my room into it and change to get ready to go home. Here's what I suggest:
1) Bring entertainment (laptop, cell phone, tablet and charger.)
2) Clothes for hubby/boyfriend
3) Clothes to wear home
Good luck!
1) Bring entertainment (laptop, cell phone, tablet and charger.)
2) Clothes for hubby/boyfriend
3) Clothes to wear home
Good luck!
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About Madison Boyer
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Current: Deland, Florida
Birth: October 17
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In a startling new discovery, revealing just how sneakily deceitful the current administration is, it appears as though sanctions have secretly been placed on American gun owners. As our Second Amendment is impeded upon on an almost daily basis, it seems as though the government is stooping to new lows in order to proceed with their unconstitutional agenda.
In an almost covert form, the Department of Justice (DOJ) has enacted what they are calling, "Operation Choke Point," right under America's nose. Effectively going around gun rights activists, under the veil of financial legislation, Holder and his goon squad passed a law making it even harder for those dealing with guns to effectively go about their business.
(See also: Obama Signs Executive Order To Legalize BLM's Land Grabs)
In a blatant anti-Second Amendment effort, the DOJ teamed up with the FDIC and the Consumer Financial Protection Bureau in order to put a stop to those complicit in "high-risk activities." Funnily enough, adding to their list of seemingly innocuous activities in which banks are urged to be wary of, and outright suggested they avoid, those selling firearms were sneakily slipped into the law.
The CEO of the Electronic Transactions Association, Jason Oxman, recently wrote on the matter saying, "Federal law enforcers are targeting merchant categories like payday lenders, ammunition and tobacco sales, and telemarketers – but not merely by pursuing those merchants directly." Furthermore he mentioned, "Rather, Operation Choke Point is flooding payments companies that provide processing service to those industries with subpoenas, civil investigative demands, and other burdensome and costly legal demands."
Effectively what the DOJ has done here is raised the cost of dealing with entities that would sufficiently fit within the list of those considered, "high-risk." Basically deeming those companies, or individuals as toxic (and now costly), the picture starts to become a little clearer.
(See also: DOJ Forcing Banks To Drop Gun Store Clients)
In an effort to put the hurt on the gun environment in any way possible, Obama's administration is, in essence, trying to suffocate those who deal in guns financially. After all, if you can't interact financially, then you really have no business at that point.
According to Oxman, banks will soon, "simply stop providing service to such merchants." The worst part in all this is that, "it's working – payments companies across the country are cutting off service to categories of merchants that – although providing a legal service – are creating the potential for significant financial and reputational harm as law enforcement publicizes its activities," Oxman mentioned.
Those managing bitcoin services have already been reported to deny those working in gun sales because of Operation Choke Point. Is it really that big of a shock however that perhaps the most politicized Attorney General in America's history has taken it upon himself to fight against guns?
Last I knew, he was supposed to be focused on matters of the law, not cultural issues. That being said, I think we can see how blind the justice is through the eyes of Mr. Holder.
One last thing really has my head spinning – the name "Operation Choke Point." Now yes, this legislation is effectively choking gun owners financially, but anyone knows that this isn't what a choke point is.
(See also: Obama Covertly Supplying Al Qaeda With Weapons)
For those of you who do not know, it's a strategic place in which the enemy is funneled with no way out and usually ends in the slaughter of whomever was forced into the choke point. Could this perhaps have some meaning regarding its futuristic intent? Now that the DOJ is funneling gun sellers in certain directions is there soon to be a slaughter?
Or is this just another instance demonstrating their severe incompetence while trying to be witty?
Let us know in a comment below.


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BOYS FROM THE WOOD A London childhood in the 1940's by John Simons Co-writer: Joan Stanley Even though it had its share of celebrities, in the 1940's there were quite a few council flats, building sites, and bombed houses punctuating the streets of expensive houses of St. Johns Wood. This was schoolboy Tommy Harris's playground. Rationing and low wages made life hard, but his working class parents were proud, and endeavoured to bring up their family to know right from wrong. With no pocket money as such, Tommy lived on his wits, finding various ways to occupy his time with his friend Tiny. Boys From The Wood is a charming, autobiographical story set in post-war Britain. It is a humorous tale, tinged with a little sadness as Tommy remembers events from the war in his dreams. This book will delight many young readers, and give them an insight into life in 1940's London. At the same time older readers can enjoy a trip back to yesteryear.
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Description/Uses/Merits of this ΣChain
A good and fast connectivity for those intending to travel from the beautiful lands of Assam to the Holiday destination Goa. Only thing is you need to change station from SDAH to HWH as SDAH don't have a train to Goa.
22512/Σ Guwahati to Vasco Da Gama 4 days chain ΣChain
created by GOA SK TO VSG~ May 31 2016 (02:12)
|Travel Time: 64h 10m 42 halts Distance: 3102 km 1 xfers
|Departs @ 22:55
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text:
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Home > Press > Harris & Harris Announces New Board of Director
Harris & Harris Group, Inc., is pleased to announce that the Company's Board of Directors, upon recommendation by its Nominating Committee, elected Mr. Richard P. Shanley to serve as a Director of the Company, effective March 12, 2007, until the 2007 Annual Meeting of Shareholders. In connection with the election of Mr. Shanley, the Board increased the number of directors from 11 to 12, with Mr. Shanley filling the vacancy created thereby. Mr. Shanley was also named to the Audit Committee, Valuation Committee and Independent Directors' Committee. There is no arrangement or understanding between Mr. Shanley and any other person pursuant to which Mr. Shanley was selected as a director of the Company. Mr. Shanley does not have any direct or indirect material interest in any existing or proposed transaction to which the Company is or may become a party.
From February 2001 to December 31, 2006, Mr. Shanley, age 59, was a partner of Deloitte & Touche LLP. From March 1976 to January 2001, he was employed by Eisner LLP and was a partner from 1982 until 2001. During his over 30 years of public accounting experience, he served as lead audit partner on numerous audit engagements for both public and private companies, including those requiring application of Sarbanes-Oxley Section 404, as well as public stock offerings, primarily for biotech, pharmaceutical and high-tech companies, including companies enabled by nanotechnology. He has been actively involved on the Biotech Council of New Jersey, the New Jersey Technology Council, the New York Biotechnology Association, the Connecticut Venture Group, the Biotechnology Industry Organization and the NanoBusiness Alliance. He is an active member of the New York State Society of Certified Public Accountants and the American Institute of Certified Public Accountants. He is currently serving his third term on the New York State Society of CPA's Professional Ethics Committee. He is a licensed Certified Public Accountant in New Jersey and New York. He was graduated from Fordham University (B.S.) and Long Island University (M.B.A. in accounting).
About Harris & Harris Group, Inc.
Harris & Harris Group is a publicly traded venture capital company that makes initial investments exclusively in tiny technology, including nanotechnology, microsystems and microelectromechanical systems (MEMS). The Company's last 31 initial private equity investments have been in tiny technology-enabled companies. The Company has 21,341,029 shares of common stock outstanding.
Detailed information about Harris & Harris Group and its holdings can be found on its website at http://www.TinyTechVC.com .
This press release may contain statements of a forward-looking nature relating to future events. These forward-looking statements are subject to the inherent uncertainties in predicting future results and conditions. These statements reflect the Company's current beliefs, and a number of important factors could cause actual results to differ materially from those expressed in this press release. Please see the Company's Registration Statement on Form N-2 and Annual Report on Form 10-K for the year ended December 31, 2005, filed with the Securities and Exchange Commission for a more detailed discussion of the risks and uncertainties associated with the Company's business, including but not limited to the risks and uncertainties associated with venture capital investing and other significant factors that could affect the Company's actual results. Except as otherwise required by Federal securities laws, Harris & Harris Group, Inc., undertakes no obligation to update or revise these forward-looking statements to reflect new events or uncertainties. The reference to the website http://www.TinyTechVC.com has been provided as a convenience, and the information contained on such website is not incorporated by reference into this press release.
For more information, please click here
Harris & Harris Group, Inc.
Charles E. Harris, 212-582-0900
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The Volkswagen brand says it has repositioned its organisation for vehicle development effective from the beginning of the year. The reorganisation reflects the company's ongoing response to its diesel emissions crisis and its review of the company's organisation - especially lines of accountability and responsibility.
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[Computational Complexity] Early or Late
Expand Messages
As you can see from the timestamp of this post, I came into work quite early this morning. I had to drive and needed an early start (about 6 AM) to beat Chicago traffic and get a good parking spot. When I arrived I saw another professor in his office. I knew he wasn't the early type and likely spent the night here. When I said "hello," he replied "deadline."
Which one of us is keeping the crazier hours?
Posted by Lance to Computational Complexity at 10/12/2005 07:23:00 AM
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Diwali is a festival of lights, a holiday celebrated in India by Hindus, Buddhists, Sikhs and Jains. The dates vary from year to year, but it falls some time in October or November. At Diwali, oil lamps are lit to symbolize good winning over evil, and to remind people of their own inner light. At night, fireworks light up the sky for the celebration. You can use this light theme to create some beautiful Diwali greeting cards to share with your friends and family during this festive season.
Things You'll Need
- Black card stock
- Scissors
- Paper in assorted colors
- Pencil
- Glue stick
- Letter stickers or die cuts
- Glitter glue pens
Mosaic Card
Cut and fold a piece of black card stock to the size card you want to make.
Collect some scrap paper or card stock in different bright colors. You can use any colors you like, but you should definitely include yellow or gold.
Cut the scrap paper into tiny pieces of confetti, approximately 1/8 inch pieces. You can make them all square, or you can cut random shapes. Keep the colors in separate piles.
Use a pencil to draw a light outline of a diya, a small, round, bowl-shaped oil burning lamp (see "Resources"). Above the diya, draw a flame in the shape of a teardrop.
Glue the tiny yellow or gold paper pieces to the flame using a glue stick. Place the pieces close together, leaving just a sliver of black showing between them.
Glue other colors of your choice the same way to decorate your lamp.
Use letter stickers or die cuts to spell out "Happy Diwali" across the top of the card.
Print your message for the inside of the card on a light-colored piece of paper. Cut it to fit the inside of the card and glue it to the black card stock.
Fireworks Card
Cut and fold a piece of black card stock.
Use a pencil to draw lines on the front of the card that resemble fireworks. Fireworks are drawn by making dots, dashes, curved or squiggly lines radiating outward from a central point. You can resize and print the fireworks coloring page in the "Resources" to use as a template, if you prefer. You can overlap two or three to make it look like a big display.
Trace over your fireworks lines with colorful glitter glue pens. If you are overlapping firework bursts, use one or two different colors per burst so they can be distinguished from one another. If you are making one big burst, make it multicolored to give it pizazz.
Write a Diwali greeting across the bottom of the card cover with your glitter glue. Allow the glue to dry.
Print your message for the inside of the card on a light-colored piece of paper. Cut it to fit the inside of the card and glue it to the black card stock.
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