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Structural and Biochemical Properties of Lipid Particles from
Receivedforpublication,January16,2008,andinrevisedform,April14,2008 Published,JBCPapersinPress,April22,2008,DOI10.1074/jbc.M800401200
Tibor Czabany‡1, Andrea Wagner‡1, Dagmar Zweytick§, Karl Lohner§, Erich Leitner¶, Elisabeth Ingolic?,
and Gu ¨nther Daum‡2
FinestructureResearch,GrazUniversityofTechnology,Steyrergasse17/III,A-8010 Graz, Austria
The two most prominent neutral lipids of the yeast Saccharo-
myces cerevisiae, triacylglycerols (TAG) and steryl esters (SE),
the two SE synthases Are1p and Are2p. In this study, we made
use of a set of triple mutants with only one of these acyltrans-
acyltransferases, LP from triple mutants contained only TAG or
SE, respectively, with specific patterns of fatty acids and sterols.
Biophysical investigations, however, revealed that individual neu-
tral lipids strongly affected the internal structure of LP. SE form
the LP. We propose that this structural arrangement of neutral
Fatty acids and sterols are important building blocks of
biomembranes. To maintain balanced cellular levels of these
components, a certain portion is put on hold in the form of
complex neutral lipids. The yeast Saccharomyces cerevisiae
similar to other eukaryotic cells stores fatty acids and sterols in
the form of triacylglycerols (TAG)3and steryl esters (SE). Both
TAG and SE are hydrophobic molecules that are not soluble in
the cytosol but are also not typical bilayer membrane lipids.
Consequently, they need to be stored in specific subcellular
lets, lipid bodies, or oil bodies. Yeast LP are small spherical
organelles with an approximate diameter of 400 nm consisting
of 95% neutral lipids, TAG, and SE, and only small amounts of
phospholipids and proteins (1). The highly hydrophobic core
layer containing a well defined set of proteins (2).
Identification of LP proteins in various types of cells has
changed the view of this organelle. The presence of certain
enzymes on the surface of LP suggests that besides lipid stor-
age LP are involved in various metabolic processes. In S.
cerevisiae ?40 proteins are known to be present on the sur-
face of LP. Strikingly, most of these proteins are enzymes
involved in lipid metabolism. As prominent examples, LP
from S. cerevisiae harbor enzymes of phosphatidic acid bio-
synthesis (3), fatty acid activation (4–6), sterol biosynthesis
(7), but also TAG biosynthesis and degradation (8–11) and
SE hydrolysis (12–14).
The biogenesis of LP is still a matter of dispute and is under
investigation in several laboratories. Whereas little is known
about the cell biology of this process, enzymes involved in the
formation of major LP components, TAG and SE, have been
identified within the last few years (for recent review see Refs.
15 and 16). In the yeast as in other types of cells, the last step of
TAG synthesis requires acylation of diacylglycerol. Two differ-
ent mechanisms of TAG synthesis have been demonstrated in
S. cerevisiae. First, the acyl-CoA-independent acyltransferase
acyl donor (17, 18) and combines functions of a phospholipase
A2(or B) and an acyltransferase. Lro1p appears to be exclu-
sively localized to the endoplasmic reticulum (ER). The
other enzyme involved in TAG synthesis is Dga1p, which is
an acyl-CoA-dependent acyltransferase. Under standard
growth conditions, Dga1p appears to be more active than
Lro1p (9). In the yeast, Dga1p is dually localized to the ER
and LP. The TAG synthase activity of Dga1p on LP demon-
strated in vitro (19) may be responsible for the formation of
TAG directly at its site of storage.
acyltransferases named Are1p and Are2p (20, 21). Both
enzymes are localized to the ER and utilize acyl-CoA as acyl
donor (7), but they differ in regulation and substrate specificity
Are2p includes most of the acyl-CoA:sterol acyltransferase
activity in the yeast and esterifies preferentially ergosterol, the
final product of sterol synthesis in the yeast (20, 22–24). Inter-
estingly, Are1p was shown to exhibit increased activity under
* This work was supported by the Fonds zur Fo ¨rderung der Wissenschaftli-
chenForschunginO¨sterreichProjects18857andW901-B05(toG. D.).The
page charges. This article must therefore be hereby marked "advertise-
1Both authors contributed equally to this work.
2To whom correspondence should be addressed: Institute of Biochemistry,
43-316-873-6462; Fax: 43-316-873-6952; E-mail: guenther.daum@
3The abbreviations used are: TAG, triacylglycerol; CE, cholesteryl ester; ER,
CHAPS, 3-[(3-cholamidopropyl)dimethylammonio]-1-propanesulfonic
acid; MES, 4-morpholineethanesulfonic acid; CER, cholesterol-rich; LDL,
chromatography; MS, mass spectrometry; DSC, differential scanning calo-
rimetry; SAXS, small angle x-ray scattering.
THE JOURNAL OF BIOLOGICAL CHEMISTRY VOL. 283, NO. 25, pp. 17065–17074, June 20, 2008
© 2008 by The American Society for Biochemistry and Molecular Biology, Inc.Printed in the U.S.A.
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hypoxic conditions and to prefer sterol precursors as sub-
strates. The presence of two SE synthases with overlapping
function, which are subject to differential regulation, might
under optimal or suboptimal conditions of ergosterol biosyn-
thesis. Screening for yeast mutants that were nonviable in the
(25). Besides their dependence on sterol esterification for
ature-sensitive, and nonviable under anaerobic conditions.
a possible role for Arv1p in sterol trafficking to the plasma
LP as a storage compartment would be useless without
enzymes that mobilize the lipid depots, especially fatty acids
and sterols from TAG or SE. In S. cerevisiae, three TAG lipases
and two SE hydrolases present on the surface of LP were iden-
tified (10–14). The major TAG lipase of S. cerevisiae identified
at the molecular level and by function is Tgl3p (10). Based on
homology to Tgl3p, two additional TAG lipases, Tgl4p and
Tgl5p, were also identified. Similar to Tgl3p, Tgl4p and Tgl5p
are involved in TAG catabolism in vivo, exhibit enzymatic
activity in vitro when purified close to homogeneity, and local-
ize to LP close to their substrate (11, 26).
Recently, three SE hydrolases of the yeast S. cerevisiae
were identified. The highest SE hydrolase activity present in
the plasma membrane (7) was attributed to Yeh2p (12, 13).
The physiological role of this SE hydrolase in the plasma
membrane is still a matter of dispute. In contrast, two other
members of the yeast SE hydrolase family, Yeh1p and Tgl1p,
were localized to LP (13, 14). These two enzymes exhibited a
very low activity in vitro, but a triple deletion in combination
with YEH2 completely abolished SE hydrolysis in the yeast.
In an hem1? deletion strain mimicking anaerobic growth,
Yeh1p became the most important SE hydrolase, whereas
Yeh2p and Tgl1p were largely inactive under this condition
Despite the progress that has been made during the last few
years to identify gene products involved in neutral lipid meta-
bolism, the biogenesis of the neutral lipid storage compart-
ment, the LP, is still poorly understood. A rather fundamental
although highly important question related to this problem is
coordination of the different enzyme activities contributing to
biosynthesis and degradation of neutral lipids. To address this
question from the viewpoint of LP formation, we studied the
contribution of individual TAG and SE synthases to this proc-
ess. For this purpose, we employed triple mutants (TM) with
only one of the four acyltransferases active, Dga1p, Lro1p,
the following: (i) the influence of neutral lipid-synthesizing
sterol synthesis; and (ii) the structure of LP variants with
unusual but well defined neutral lipid composition. The result
of this study is a novel model of yeast LP structure presented
of this organelle.
Yeast strains used throughout this study are listed in the Table
1. Cells were grown aerobically to the late logarithmic (16 h) or
stationary phase (24 h) at 30 °C in YPD medium containing 1%
yeast extract (Oxoid), 2% peptone (Oxoid), and 2% glucose
(Merck). Media were inoculated with precultures grown aero-
bically for 24 h in YPD medium at 30 °C to an A600of 0.1.
The yeast lipid particle (LP) fraction was obtained at high
purity from cells grown to the early stationary phase as
described by Leber et al. (1). The isolation of other subcellular
fractions used in this study was described by Zinser and Daum
Western blot analysis (see below).
Lowry et al. (29) using bovine serum albumin as a standard.
lizedin0.1%SDS,0.1 MNaOHpriortoquantification.WhenLP
fractions were analyzed, samples were delipidated prior to pro-
tein quantification. For this purpose, nonpolar lipids were
was withdrawn; residual diethyl ether was removed under a
SDS-PAGE was carried out by the method of Laemmli (30)
using10%SDSgels.Samplesweredenaturatedat37 °Ctoavoid
hydrolysis of polypeptides. Proteins on gels were detected by
staining with Coomassie Blue. Western blot analysis was per-
ies used in this study were from rabbits and were directed
against Erg1p, Por1p, Wbp1p, and Prc1p/CPY. Peroxidase-
conjugated secondary antibody and enhanced chemilumines-
cent signal detection reagents (SuperSignalTM, Pierce) were
used to visualize immunoreactive bands.
Lipid Extraction and Analysis—Lipids from yeast cells were
extracted as described by Folch et al. (32). For quantification of
neutral lipids, extracts were applied to Silica Gel 60 plates with
the aid of a sample applicator (CAMAG, Automatic TLC Sam-
pler 4, Muttenz, Switzerland), and chromatograms were devel-
oped in an ascending manner by a two-step developing system.
ume) was used as mobile phase, and plates were developed to
further developed to the top of the plate using the second
mobile phase consisting of light petroleum:diethyl ether (49:1;
water, 60 ml of methanol, and 4 ml of concentrated sulfuric
acid, briefly dried, and heated at 100 °C for 30 min. The lipids
were then quantified by densitometric scanning at 400 nm
using a Shimadzu dual-wave length chromatoscanner CS-930
with triolein and cholesteryl ester as standards.
Fatty acids were analyzed by gas-liquid chromatography
(GLC). Lipid extracts prepared as described above were sub-
jected to methanolysis using BF3/methanol and converted to
methyl esters as described by Morrison and Smith (33). Fatty
acid methyl esters were separated using a Hewlett-Packard
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illary column (15 m ? 0.25 mm inner diameter ? 0.50 ?m film
thickness) and helium as carrier gas. Fatty acids were identified
by comparison to commercial fatty acid methyl ester standards
(NuCheck, Inc., Elysian, MN).
Individual sterols from whole cells or LP were identified and
quantified by GLC/mass spectrometry (MS) after alkaline
hydrolysis of lipid extracts (34). GLC/MS was performed on a
Hewlett-Packard 5890 gas chromatograph equipped with a
mass selective detector (HP 5972), using an HP5-MS capillary
column (30 m ? 0.25 mm ? 0.25 ?m film thickness). Aliquots
of 1 ?l were injected in the splitless mode at 270 °C injection
temperature with helium as a carrier gas at a flow rate of 0.9
gram was used: 1 min at 100 °C, 10 °C/min to 250 °C, and 3 °C/
min to 310 °C. Mass spectra were acquired in scan mode (scan
range 200–550 atomic mass units) with 3.27 scans/s. Sterols
were identified based on their mass fragmentation pattern.
Enzymatic Assays—For the preparation of [14C]palmitic
acid-labeled phospholipids, cells were inoculated to an A600of
[14C]palmitic acid (PerkinElmer Life Sciences) and incubated
for 4 h at 30 °C. The cells were then harvested, and lipids were
extracted and separated as described above. The phospholipid
methanol (2:1, v/v), and radioactivity was measured by liquid
scintillation counting using LSC safety mixture (Baker,
Deventer, The Netherlands) and 5% water as a scintillation
The acyl-CoA:diacylglycerol acyltransferase assay was per-
formed in a final volume of 100 ?l containing 6 nmol of
[14C]oleoyl-CoA (80,000 dpm), 0.025 mM dioleoylglycerol, 0.5
mM CHAPS, 150 mM Tris-Cl (pH 7.0), 15 mM KCl, 15 mM
lipid:diacylglycerol acyltransferase assay was performed in the
final volume of 100 ?l containing [14C]palmitic acid-labeled
phospholipids (70,000 dpm, 188 nmol), 0.05 mM dioleoylglyc-
erol, 150 mM Tris-Cl (pH 7.0), 15 mM KCl, 15 mM MgCl2, and
assay was performed in the final volume of 100 ?l containing 6
nmol of [14C]oleoyl-CoA (80,000 dpm), 0.025 mM ergosterol,
and 100 ?g of protein (35).
Incubations were carried out for 20 or 30 min at 30 °C and
terminated by addition of 300 ?l of chloroform:methanol (2:1,
v/v). Lipids were extracted twice for 10 min with shaking using
300 ?l of chloroform:methanol (2:1, v/v). Organic phases were
combined and washed twice using methanol:water:chloroform
(47:48:3; per volume). The organic phase was taken to dryness
under a stream of nitrogen. Lipids were dissolved in 30 ?l of
chloroform:methanol (2:1, v/v), separated by TLC as described
above, and visualized on TLC plates by staining with iodine
vapor. Bands were scraped off, and radioactivity was measured
by liquid scintillation counting using LSC safety mixture
(Baker, Deventer, The Netherlands) and 5% water as a scintil-
lation mixture.
Electron Microscopy of Whole Yeast Cells—Yeast cells grown
on YPD were sedimented by centrifugation, rinsed in 0.1 M
cacodylate buffer (pH 7.4), and fixed with 3% glutaraldehyde in
0.1 M cacodylate buffer (pH 7.4) containing 5 mM CaCl2and 5
were then washed twice with 0.1 M cacodylate buffer (pH 7.4)
and postfixed in 4% KMnO4, prepared in distilled H2O for 90
min. After rinsing the pellet four times, cells were treated with
0.5% sodium metaperiodate for 15 min at room temperature,
washed once more with distilled H2O, and dispersed in a 2%
agarose solution (1:1). The solidified agar containing the cells
was cut into small cubes of about 1 mm3and placed into 2%
ice. Blocks were washed with distilled H2O and dehydrated
through a graded series of acetone, 10 min each, followed by
three washes with 100% acetone for 15 min each. Samples
were incubated in 2:1, 1:1, and 1:2 mixtures of acetone/Spurr
resin for 2 h, respectively. The last incubation step was per-
removal of the Spurr resin, agar cubes were incubated in
fresh 100% Spurr resin for another 2 h. The samples were
then placed into gelatin capsules containing fresh Spurr
resin (100%), and capsules were placed in a polymerization
oven at 70 °C for 48 h (12). Using a diamond knife (Diatome),
75-nm ultrathin sections were cut on a Leica Ultracut E
ultramicrotome and collected into 75 mesh copper palla-
dium grids with a support film of pioloform. Sections were
stained in 1% uranyl acetate in distilled H2O for 20 min,
washed, and stained once more in lead citrate (36). Images
were produced on a Tecnai G212 (FEI Co.) equipped with a
CCD camera (Gatan Bioscan) at 100 kV.
Differential Scanning Calorimetry (DSC)—Calorimetric
experiments were performed with a differential scanning
calorimeter (VP-DSC) from MicroCal, Inc., Northampton,
MA. Samples of LP prepared from wild type and mutant
strains were used undiluted. A buffer containing 0.25 M sor-
bitol, 10 mM MES/Tris, and 0.2 mM Na2EDTA (pH 6.9) was
used to fill the reference cell, and samples were degassed
before measuring. Heating scans were performed at a scan
rate of 15 °C/h between 1 and 65 °C. Pre-scan thermostating
was allowed for 60 min. Enthalpies were calculated by inte-
grating the peak area after base-line adjustment and normal-
ization to SE concentrations using the MicroCal Origin soft-
ware (VP-DSC version).
SAXS—X-ray diffractograms were recorded with a SWAX-
camera (HECUS x-ray Systems, Graz, Austria) that was
mounted on a sealed tube Seifert generator (Ahrensburg, Ger-
many) operating at 2 kilowatts (50 kV and 40 mA). The x-ray
beam was filtered for CuK?-radiation (? ? 1.54 Å) using a
nickel foil and a pulse-height discriminator built into the
detection system. The scattered intensity was recorded with
two linear position-sensitive detectors (PSD 50, HECUS
X-ray Systems, Graz, Austria) simultaneously in the small
angle regimes of 10?3Å?1? q ? 1 Å?1(SAXS), where q ? 4
? sin(?)/? is the modulus of the scattering wave vector. Cal-
ibration was performed with silver stearate. Samples of LP
prepared from yeast wild type and mutant strains were
loaded into thin walled quartz capillaries thermally equili-
brated for 60 min at 1 °C before initiating data acquisition.
Temperature was controlled with an accuracy of 0.1 °C with
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a programmable Peltier unit. Diffraction patterns were
recorded at each temperature for 7200 s. Measurements
were performed at 1, 35, and 65 °C.
to determine the mean diameter and size distribution of LP
from yeast wild type and mutant strains using a Zetasizer
3000HS (Malvern Instruments, Worcestershire, UK). The
instrument was equipped with a 10-milliwatt helium-neon
culation was derived from an autocorrelation function. For
measurements, samples were diluted with water that was fil-
tered (pore size 0.02 ?m) prior to use to avoid dust particles
interfering with measurements. To check reproducibility, a
series of three measurements were performed at room
S. cerevisiae produces TAG and SE by catalysis of four acyl-
transferases, Dga1p, Lro1p, Are1p, and Are2p. To address the
neutral lipids and thus to the biogenesis of LP, we used triple
mutants (TM) with only one of the four acyltransferases active
as an experimental system. LP from the different TM were iso-
lated at highest purity and subjected to various biochemical,
cell biological, and biophysical analyses.
Neutral Lipids of Triple Mutants—Neutral lipids of the yeast
start accumulating at the end of the logarithmic phase and
reach a high level in the stationary phase when cells stop divid-
ing. Therefore, the correct time point of harvesting was impor-
tant to compare the amounts of neutral lipids from wild type
and TM. Because wild type and TM used in this study (Table
mid-stationary phase (24 h). As can be seen from Fig. 1A, the
amount of TAG from wild type W303 increased approxi-
mately three times between the two time points taken. A TM
with only Lro1p active (TM LRO1?) showed a marked
decrease of the TAG amount compared with wild type after
16 or 24 h of growth, respectively. However, the level of TAG
was also increased in this strain from the late logarithmic to
that Lrop1p in contrast to Dga1p was more active in loga-
rithmic phase (9). A triple mutant with Dga1p as the only
active acyltransferase (TM DGA1?) exhibited a TAG level
that was similar to wild type, showing a similar increase in
the TAG level from 16 to 24 h of growth. Dga1p had been
assumed to remain active within the stationary phase (9). It
has to be mentioned that neither TM DGA1?nor TM
LRO1?contained SE.
TM with only Are1p or Are2p active (TM ARE1?and TM
ARE2?) form SE as the major neutral lipid components (Fig.
1B). Using sensitive radiolabeling techniques, it had been
shown before that Are1p and Are2p are able to synthesize also
minor amounts of TAG (9, 37). In our experiments, however,
TAG was not detectable in these strains because of the less
sensitive analytical method employed in this study. In the wild
type, the amount of SE is increased by 60% from 16 to 24 h of
growth. This enhancement appears to be attributable to both
Are1p and Are2p, because SE in TM ARE1?and TM ARE2?
increased accordingly over the observed time period. The con-
only one active acyltransferase from cells grown to the late logarithmic
ent experiments. The amounts of TAG and SE per A600found in the wild type
after 24 h were set at 100%. A, relative amount of TAG from wild type, TM
DGA1?, and TM LRO1?. B, relative amount of SE from wild type, TM ARE1?,
and TM ARE2?.
Yeast strains used in this study
W303 (wild type)
aQM indicates quadruple mutant.
Relevant genotypeSource of Ref.
SUC2 GAL?mal mel ade2-1 can1-100 his3-11,15 leu2-3 112 trp1-1 ura3-1
W303; MAT? lro1::TRP1 are1::HIS3 are2::LEU2 ADE2 met ura3
W303; MAT? dga1::KanMX4 are1::HIS3 are2::LEU2 ADE2 met trp1 ura3
W303; MAT? dga1::KanMX4 lro1::TRP1 are2::LEU2 ADE2 met his3 ura3
W303; MAT? dga1::KanMX4 lro1::TRP1 are1::HIS3 ADE2 met leu2 ura3
W303; MAT? dga1::KanMX4 lro1::TRP1 are1::HIS3 are2::LEU2 ADE2 met ura3
ScanBi Ltd., Alnarp, Sweden (37)
ScanBi Ltd., Alnarp, Sweden
ScanBi Ltd., Alnarp, Sweden
ScanBi Ltd., Alnarp, Sweden
ScanBi Ltd., Alnarp, Sweden
ScanBi Ltd., Alnarp, Sweden
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compared with Are2p confirming previous results by others
(20, 24, 37).
Interestingly, enzyme activities in vitro of the four different
acyltransferases measured did not completely reflect the levels
with the same substrate in the same assay and exhibited the
Lro1p were measured with different substrates and different
assays and therefore could not be directly compared. Both
enzyme activities were markedly decreased in mutant strains
(Fig. 2). We have to take into account, however, that enzyme
vivo, but we also have to consider
that changed cellular conditions in
TM may be responsible for the
observed alterations of the enzyme
Neutral Lipid Product Profile
Generated by Individual Acyl-
transferases—One explanation for
the existence of TAG and SE syn-
thases with overlapping function
may be different substrate specific-
ity and consequently formation of
different product
fatty acid and sterol analysis of TAG and SE formed by the
respective individual acyltransferases.
The fatty acid composition of total lipids from the four dif-
not shown). Thus, the overall cellular fatty acid formation was
not affected by the absence of three neutral lipid-synthesizing
and SE from the different TM varied compared with wild type.
As can be seen from Fig. 3A, both the TM DGA1?and TM
LRO1?strain contained more unsaturated fatty acids in the
to a shift from C-16:0 to C-16:1, but less from C-18:0 to C-18:1
species. These changes resulted in a marked alteration of the
ratio of unsaturated to total fatty acids in TAG from TM
DGA1?and TM LRO1?compared with wild type (Table 2).
The ratio of C-16 to C-18 fatty acids was increased in TM
DGA1?but unchanged in TM LRO1?.
Fatty acid analysis of the SE fractions revealed that in TM
ARE1?the amount of unsaturated fatty acids was slightly
decreased compared with wild type (Fig. 3B and Table 2). No
such effect was observed in the TM ARE2?strain, confirming
type and slightly increased in TM ARE2?.
Similar to a certain specificity of neutral lipid synthesizing
acyltransferases for fatty acids as substrates, Are1p and Are2p
exhibited some preferences for sterol moieties incorporated
are the four most abundant esterified sterols stored in the form
sterol pattern of SE from TM ARE2?was almost identical to
wild type, Are2p showed a slight preference for sterol precur-
sors over ergosterol compared with wild type confirming pre-
vious results by Zweytick et al. (22) and Valachovic et al. (38).
regulatory effect of neutral lipid storage on sterol biosynthesis
(Table 3). TM DGA1?and TM LRO1?, which lacked sterol
esterification, accumulated ergosta-7,22-dienol and ergosta-7-
enol at the expense of other sterol precursors. This effect may
be explained by down-regulation of Erg3p as reported previ-
ously by Arthington-Skaggs et al. (39) and will be discussed
pholipid:diacylglycerol acyltransferase (B) was measured using crude ER fractions. Strains are described in
as 100%. The quadruple mutant (QM) dga1?lro1?are1?are2? was used as negative control.
FIGURE 3. Fatty acid analysis of neutral lipids. A, fatty acid analysis of TAG
bars), and TM ARE2?(gray bars) strains. Values are shown as percent of total
fatty acids and are from three independent measurements with the respec-
acid; C18:1, oleic acid.
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U.S. lawmakers and industry advocates need to do more to address a "critical" shortage of skilled truck technicians and mechanics, according to Bill Graves, CEO of American Trucking Associations.
The planned Nov. 2 rate hike will be FedEx's second in nine months, according to multiple media reports.
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AndroidGals Seeks Contributors
ROSE Blog: Rikki's Open Source Exchange
In the latest issue of Linux Pro Magazine, I wrote about mobile app development startups and the new job opportunities around mobile technology. I also mentioned that we're accepting article proposals for our new magazine, Smart Developer.
The AndroidGals website is also looking for contributors: WANTED: Females who have an opinion on Android! I certainly don't have time to contribute because I'm much too busy playing WordFeud with strangers on my HTC Incredible, but maybe you have more bandwidth than I do. (I also love my RunKeeper app and couldn't get anywhere without Google Maps on my phone.)
Happy Androiding!comments powered by Disqus
Issue 180/2015
Buy this issue as a PDF
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The deal strengthens the bond between the filmmakers and Searchlight, which picked up MMMM at Sundance and is releasing it now in theatres.
FILM REVIEW: Martha Marcy May Marlene
Borderline is comprised of Antonio Campos, Sean Durkin and Josh Mond. Durkin wrote the script for MMMM and made his directorial debut on the drama. Campos and Mond acted as producers along with Patrick Cunningham and Chris Maybach. MMMM, starring Elizabeth Olsen, won the dramatic feature directing award at Sundance and went on to unspool at the Festival de Cannes, Toronto International Film Festival and New York Film Festival among others. Olsen has generated some Oscar buzz for her breakout performance.
The principals call Searchlight "an ideal home" and say the move takes their company to a new level.
AUDIO: Sean Durkin Discusses 'Martha Marcy May Marlene's' Rare Journey Through Four Major Film Fests
The latter is certainly true, as it will give the company—created in 2003 and having produced several acclaimed shorts, music videos and features—a kind of stability almost impossible for indie filmmakers to find.
The deal also should serve to calm one of the biggest worries that indie filmmaker have—wondering what its next distribution outlet will be.
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10 top natural attractions in the Karoo
What the Outback is to Australia, the Karoo is to South Africa.
It occupies 400 000 km²- practically one-third of the country's total area; a vast, semi-arid region in the heart of the country, bounded by mountains in the south and west, and the huge Senqu, or Orange River, on its eastern and northern flanks.
This area of little rainfall, blue and cloudless skies, and extreme temperatures may have acted as a barrier to the interior from Cape Town for early settlers, but today it has come out of isolation to become a major tourist attraction – its immense spaces, incredible mountains, idiosyncratic towns and promise of escape the reason for its popularity.
These 10 top natural attractions are only a taste of what the Karoo holds in store for you. Continued


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David Duchovny, Robin Williams, Tea Leoni. An American living in Paris comes to terms with himself and his family in this touching story written and directed by Duchovny. 2004/color/97 min/PG-13.
House of D is a bittersweet, moving story of an American expatriate's painful decision to come to terms with the childhood he fled in early 1970s New York City. David Duchovny wrote and directed this comedy-drama; he also stars as the adult version of the film's hero, Tom Warshaw, an illustrator who has spent most of his life in Paris and decideson the occasion of his son's birthdayto finally reveal long-withheld facts about his past.
The bulk of the story, told in flashback, portrays 13-year-old Tom (Anton Yelchin) as a quick-witted prince of his neighborhood, a delivery boy who knows every eccentric on his bicycle route and a Catholic school kid fond of playing pranks on his clueless French teacher and soulful principal (Frank Langella). His best friend is the school's mildly retarded, 41-year-old janitor, Pappas (Robin Williams), and his advisor on matters of the heart is Lady (Erykah Badu), a prison inmate whom the fatherless Tom (or Tommy, as he's called in 1973) can neither see nor touch. Tommy's vivacity is an asset at home, where his mother (Tea Leoni), a grieving widow with a mounting addiction to pills, is slipping away from her son's ability to help. Duchovny's screenplay sometimes borders on the precious: A number of scenes are enamored with their own boldness and originality, as if Duchovny has been squirreling away lots of colorfully expressive storytelling details for years, and unloaded them here. But that flaw all but disappears in the glow of House of D's emotional resonance and honesty, not to mention several exceptional performances. Among these is Zelda Williams's work as Tommy's sage-beyond-her-years girlfriend, Melissa, whose name offers a suitable excuse to work a rather lovely Allman Brothers song into the soundtrack. --Tom Keogh
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Cute movie, I wish more people had heard of it. Lots of funny moments and some sombering ones too.Published 2 months ago by whit893
For one of Williams' last projects, he's performing a very challenging role. The kid is fantastic.Published 3 months ago by abrecken
This is a great movie with a superb cast; it moved me to tears at the end. Great job of acting and directing for David Duchovny!Published 3 months ago by Stephanie Specht
This movie was one of a kind. I thought it was a very touching, coming out of boyhood into manhood movie that dealt with issues such as emotional distress from mother, to childhood... Read morePublished 4 months ago by kellee
This is such a touching story of love and disfunction. A coming of age is such hard circumstances. David Duchavny and Robin Williams were amazing in this beautiful storyPublished 4 months ago by katonahottinroof


text:
I am taking command of the fleet.
View Single Post
May 1 2013, 09:19 PM
C.E. Evans
Vice Admiral
Location: Ferguson, Missouri, USA
Re: I am taking command of the fleet.
Marsden wrote:
I find it incredible they can find that many ships to fight an invader when usually there is only one ship in range to save Earth.
I think during the TOS era, they were going with the idea that the Starfleet was fairly small and generally thinly stretched out, with starships being deployed more like today's aircraft carriers.
"Don't sweat the small stuff--it makes you small-minded..."
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text:
I see the nail! Hammer and nail. Cool!
It's Hammer Time..!!
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text:
The Statutory Accounting Principles (E) Working Group is responsible for developing and adopting substantive and nonsubstantive revisions to the Statements of Statutory Accounting Principles (SSAPs). Statutory accounting principles provide the basis for insurers to prepare financial statements for financial regulation purposes, and SSAPs are considered level 1 (highest authority) in the statutory accounting hierarchy.
Substantive statutory accounting revisions introduce original or modified accounting principles. Substantive revisions can be reflected in an existing SSAP or a new SSAP. Nonsubstantive statutory accounting revisions are characterized as language clarifications that do not modify the original intent of an SSAP.
Items for consideration by the Working Group can be submitted via a SAPWG Maintenance Agenda Submission Form (Form A) to the NAIC staff support listed on this web page.
The Emerging Accounting Issues (E) Working Group is responsible for responding to questions on the application, interpretation and clarification of statutory accounting principles. Additional information on this Working Group can be obtained via the following link:
2015 Charges
The following Subgroup reports to the Statutory Accounting Principles Working Group:
Exposure Drafts Outstanding
Statutory Accounting Principles Working Group
Last Updated August 18, 2015
Please address comment letters to Dale Bruggeman, Chair of the Statutory Accounting Principles (E) Working Group, and direct to email@example.com, firstname.lastname@example.com and email@example.com.
Ref No.
Issue Paper/ SSAP No./ Appendix
EXPOSURES WITH COMMENT DEADLINE OF Oct. 16, 2015
SSAP No. 26
Exposure: Questions Raised on BlackRock's "Calculated" Amortized Cost Valuation Proposal
Blackrock Comment Letter
(May 26, 2015)
Inquiries on the BlackRock proposal may be sent to Katie Garvey:
firstname.lastname@example.com. However, all comments on the questions / discussion items included in the exposure document shall be submitted to NAIC SAPWG staff.
The Working Group exposed questions raised in contemplation of evaluating BlackRock's "calculated" amortized cost valuation proposal.
The Working Group directed NAIC staff to work with BlackRock and others to obtain information regarding these items and draft tentative responses. (The BlackRock Comment Letter, which includes the proposal, is provided as a supplement to the exposure.) In addition to the vendor and industry questions, which have been combined in the exposure document, three discussion questions from NAIC staff are detailed on pages 4-5.
EXPOSURES WITH COMMENT DEADLINE OF Oct. 2, 2015
Issue Paper No. 151
Revisions to Measurement Method for Surplus Notes
Substantive revisions to the measurement method for holders of non-rated surplus notes and surplus notes with designations below NAIC 1. Requests comments on specific discussion points, including whether the use of amortized cost should be expanded.
SSAP No. 26
Prepayment Penalties and Amortization on Callable Bonds
Nonsubstantive revisions to clarify the yield-to-worst concept for callable bonds.
SSAP No. 97
Nonadmitted Assets and Application of the SAP Guidance
Nonsubstantive revisions to: 1) clarify accounting for non-insurance SCAs; 2) add disclosure of permitted or prescribed practices for insurance SCAs; and 3) clarify adjustments for non-insurance SCAs meeting the revenue and activity test.
SSAP No. 15
ASU 2015-03: Simplifying the Presentation of Debt Issuance Cost
Nonsubstantive revisions to reject ASU 2015-03 and maintain the current guidance for debt issuance costs.
SSAP No. 92
SSAP No. 102
ASU 2015-04: Practical Expedient for the Measurement Date of An Employer's Defined Benefit Obligation and Plan Assets
Nonsubstantive revisions to include guidance regarding interim re-measurement of plan assets and benefit obligations due to a significant event and adopt, with modification, ASU 2015-04.
SSAP No. 26 and SSAP No. 43R
Measurement Method for NAIC 5 Designations
Nonsubstantive revisions to require investments with an NAIC 5 designation held by a reporting entity that maintains an AVR to be reported at the lower of amortized cost or fair value.
Appendix F
Policy Statement Revisions – Membership and Roles of SAPWG
Revisions propose to disband the Emerging Accounting Issues (E) Working Group, with the Statutory Accounting Principles (E) Working Group incorporating a new process to issue interpretations and to increase its membership by two. Additionally, other proposed revisions reflect changes to reflect current SAPWG processes.
SSAP No. 55
Clarification of Accounting Treatment for Fees Incurred for Salvage/Subrogation Recoveries
Nonsubstantive revisions to clarify that fees incurred for salvage and subrogation recoveries shall be reported gross, regardless of whether the fees are paid to third parties or processed internally.
Prepayment Penalties and Presentation of Callable Bonds
Exposed three options for the accounting and reporting treatment of prepayment penalties: 1) investment income, 2) realized capital gains, subject to IMR and 3) realized capital gains, excluded from IMR.
SSAP No. 23
Foreign Currency Translation for Canadian Insurance Operations
Nonsubstantive revisions to clarify the optional accounting treatment for translation of Canadian Insurance Operations.
SSAP No. 97
Inclusion of Filing Guidance
Nonsubstantive revisions to incorporate a new disclosure and incorporate the filing process for SCA investments from the Purposes and Procedures Manual of the NAIC Investment Analysis Office.
ASU 2011-10: Derecognition of in Substance Real Estate
Nonsubstantive revisions to reject ASU 2011-10.
SSAP No. 1
Quarterly Reporting of Investment Schedules
Exposed agenda item with a request for quarterly investment proposals identifying what should be captured, and the form of such submissions, that considers both cost concerns and regulatory benefits.
Appendix A
Revisions to the Accounting Practices & Procedures Manual Preamble
Nonsubstantive revisions to the Preamble to: 1) restructure to clarify historical guidance, 2) include the preamble within level 4 of the SAP Hierarchy, and 3) add a narrative reference to INT 00-20.
SSAP No. 55
Title Note 25 and SSAP No. 55
Nonsubstantive revisions to provide title specific language to the quarterly disclosure of changes to reserve and loss adjusting expense liabilities for events covered in prior years.
SSAP No. 107
SSAP No. 107 Revisions – Premium Adjustments Subject to Redetermination
Nonsubstantive revisions regarding contracts subject to redetermination, which are consistent with recent revisions adopted in SSAP No. 54.
Various SSAPs
Non-Recourse Charity Notes
Substantive concept exposure requests comments on the structure of transactions that could be considered in-scope and proposals to clarify the accounting and reporting.
Application of Equity Method
Nonsubstantive revisions to clarify the guidance in paragraphs 10-12 related to application of the equity method.
SSAP No. 78
Aging and Revenue Recognition of Multi-Peril Crop policies
Nonsubstantive concept exposure to collect information regarding any needed updates to SSAP No. 78 and the intent to develop clarifying language regarding: 1) the use of the billing date for application of the 90-day rule; 2) defining the processing date or updating the term; 3) providing more specificity regarding the period of risk for purposes of earning revenue; and 4) developing a glossary of terms including dates that are specific to the federal crop programs.
Insurance-Linked Securities Disclosure
Nonsubstantive concept exposure to collect information regarding the use of insurance-linked securities by both insurers and reinsurers, with an example template of what may be requested for disclosure.
SSAP No. 65
Proposed Disclosure Regarding High Deductible Policies
Nonsubstantive revisions to capture information regarding recoverables from policyholders under high deductible policies.
SSAP No. 61R
Variable Annuity Captive Disclosures
Nonsubstantive revisions to capture information regarding reinsurance agreements with affiliated captive reinsurers.
ASU 2015-09—Financial Services – Insurance, Disclosures about Short-Duration Contracts:
Exposed agenda item requesting comments on the GAAP disclosures for short-duration insurance contracts and whether revisions to statutory accounting disclosures should be considered
MA Review
This Memo reviews older agenda items with recommendations to either dispose the agenda item without revisions to SAP or to reject as not applicable.
Dispose: 2001-21, 2003-01, 2004-17, and 2008-21.
Reject as N/A: 2007-25
*Please forward all comment letters to email@example.com, firstname.lastname@example.com and email@example.com.
conference calls & interim
meetings are available on the NAIC
Conference Call Calendar. See NAIC's revised policy statement on open meetings.
Conference Call
Monday, Oct. 19, 2015
3:00 p.m. Eastern/2:00 pm. Central/1:00 p.m. Mountain/12:00 noon Pacific/11:00 a.m. Alaska/9:00 a.m. Hawaii
Call duration: 1 hour
Introduction to
Statutory Accounting Principles
Two-Week Online Self-Study Course
Accounting Practices and Procedures Manual Updates
Maintenance Agenda
Updated September 2015
SAP to Annual Statement Disclosure Checklist Updated
for 2014 Annual Statements
Please note, in an effort to go green, the spreadsheets are spaced to view the entire width on a computer screen. At other percentages or if printing, some detail may be lost.
Policy Statements
See Appendix F – Policy Statements of the Accounting Practices and Procedures Manual for a complete listing.
Accounting Practices and Procedures Manual
States' Prescribed Differences from NAIC Statutory Accounting Principles


text:
The collected poems of Dylan Thomas
New Directions, 1953 - Poetry - 203 pages
The prologue in verse, written for this collected edition of my poems, is intended as an address to my readev3 the strangers. This book contains most of the poems I have written, and all, up to the present yea2, that I wish to preserve. Some of them I have revised a little. Preface.
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Dylan Thomas's imagery gives me goosebumps. - Goodreads
From Dylan Thomas's introduction to this v - Goodreads
Much religious imagery. - Goodreads
Review: Collected PoemsUser Review - Rex Runt - Goodreads
A brilliant introduction to the fantastic poetry of Dylan Thomas. Many great poems that you will want to read again and again. I will treasure this book Read full review
Review: Collected PoemsUser Review - Ali - Goodreads
Though wise men at their end know dark is right, Because their words had forked no lightning they Do not go gentle into that good night. . . Rage, rage against the dying of the light Read full review
When like a running grave
My world is pyramid
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bell billhook birds blessed blind blood blows bone boys boys of summer bread breaks breast breath bride burning child climb cloud curlew dark dead death DEATHS AND ENTRANCES drowned dumb dust dying DYLAN THOMAS earth eunuch eyes fall father fire fishes flame flesh flower flying garden ghost golden grave green grief gulls hair hand head hear heart heaven heron hill holy kiss land light locks loud love's lover manshape marrow mermen moon morning mountain mouth naked night poem praise pray prayer Rage rain ride rock round sail salt sand seed shade shape shell sing skull sleep snow snow blind spring stars stone sucked summer tears tell tide tongue tower town trees turn veins VERNON WATKINS voice walk wave weather wind WIND CHILLS wind on fire wings winter's tale womb wood world Turns worm wound


text:
I think Bibi is being fed the kind of information he wants to hear. Same thing happened to Bush and Nixon here in the USA. Everybody only pays attention to the news that supports what they already believe. My sense is that Americans are sick to death about hearing and dealing with the Israeli/Palestinian problem. Their opinion of Israel is NOT very positive. Maybe Bibi should examine exactly how the questions in the polls are being worded. Poll results are very easy to manipulate by just making subtle changes to the wording of the questions. This is general knowledge here.
Two seriously wounded in crime-related shooting in Netanya (Haaretz)


text:
Hôtel du Parc, Saumur
- Free WiFi and free parking
Main amenities
- 41 guestrooms
- Restaurant and bar/lounge
- Breakfast available
- Air conditioning
- Garden
- Meeting room
- Luggage storage
- Limited hour front desk
What's around
- Chateau de Saumur (2.2 mi / 3.6 km)
- Maison de Vin (1.9 mi / 3 km)
- Wine Cellar Louis de Grenelle (2.6 mi / 4.2 km)
- Notre-Dame des Ardilliers (2.6 mi / 4.2 km)
Waitress dropped our camera and we needed to drive 75 miles to replace it. It was an…
Hotel was excellent. When visiting castle there is free parking along the river
Enter dates to check prices
Hotel Info: 866-539-8117from $82
- Family Room
- Comfort Twin Room
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- Superior Twin Room
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Hotel size
- This hotel has 41 rooms
- This hotel is arranged over 2 floors
- Check-in time 2 PM-11 PM
- Check-out time is 11 AM
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If you are planning to arrive after 11 PM please contact the property in advance using the information on the booking confirmation.
Required at check in
- Credit card or cash deposit required
- Government-issued photo ID required
Travelling with others
- Pets allowed *
- Specific rooms only *
- Free WiFi in public areas
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Nearby Three Star Saumur hotels
In the hotel
- Buffet breakfast daily (surcharge)
- Restaurant
- Bar/lounge
- Bicycle rentals nearby
- One meeting room
- Front desk (limited hours)
- Luggage storage
- Number of buildings/towers - 1
- Garden
- In-room accessibility
In the room
- Air conditioning
- Iron/ironing board (on request)
- Private bathroom
- Bathtub or shower
- Hair dryer
- Satellite TV channels
- Desk
- Free WiFi
- Phone
Top Saumur hotels
Recently reviewed hotels in Saumur
- Hostellerie GargantuaFrom
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Nearby hotels
Special features
DiningDu Parc - This restaurant specializes in regional cuisine and serves breakfast and dinner. A children's menu is available. Open select days.
Hôtel du Parc, Saumur's small print
Also known as
- du Parc Saumur
- H⌠tel du Parc Saumur
Pet-friendly rooms can be requested by contacting the property at the number on the booking confirmation.
Optional extras
Rollaway beds are available for EUR 15.00 per night
Buffet breakfast is offered for an extra charge of EUR 9.50 per person (approximately)
Pets are allowed for an extra charge of EUR 6.00 per pet, per night
We have included all charges provided to us by this hotel. However, charges can vary, for example, based on length of stay or the room you book.


text:
Contact: Michael Marden
377 Commercial St
Rockport, ME 04856
Phone: 207-236-8879
Hours: 7:30-4:30 (open thru lunch)
Email: firstname.lastname@example.com
Dr. Michael E. Marden was born and raised locally. He graduated from Belfast High School, received his Bachelors Degree from the University of Maine at Orono, and his Doctor of Optometry Degree from The Massachusetts College of Optometry. He has three grown sons and lives with his wife, Ramona, in Morrill, Maine. He has examined patients' eyes in the mid-coast for over 30 years. Dr. Marden's practice focus is primary eye care for both children and adults. He offers emergency care evenings and weekends. His hobbies include cutting firewood with his brother in the winter months and umpiring youth baseball and softball in the summer months.
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11 Swan Lake Ave
Belfast Maine 04915
Hours: 8:00 - 4:00 Closed Fridays
Email: email@example.com
Phone: 207-338-1900
Fax: 338-1946


text:
Beckhams deny adoption rumors
A representative for DAVID BECKHAM and his wife Victoria has denied the couple is planning to adopt a child.
The soccer ace and his singer-turned-fashion designer wife already have a brood of four after daughter Harper, who was born last July (11), added a girl to their three boys; Brooklyn, Romeo and Cruz.
Reports published in an Australian magazine suggested the couple was desperate to add another tot to the family, and were considering giving an orphan a home.
But a spokesperson for the Beckhams has refuted the claims, telling Britain's Daily Mirror, "David and Victoria have huge respect for people that choose to adopt but this is untrue. They continue to focus on raising their own four children."


text:
Middlesbrough provides an excellent variety of places to eat out. From great places to grab a coffee, tea or light meal from the inspired vintage Olde Young Tea House to Mink, where you can take your seat on a repurposed fairground waltzer!
Middlesbrough also offers a world of restaurants to suit every palate from Italy to India, China and Portugal.
When what you really want is hearty, comfort food in a relaxed atmosphere we have you covered.
Serving delicious illy coffee, a wide variety of beverages, sandwiches and hand made snacks.
Unique Teesside cafe for foodies who love good, honest, healthy cooking.
Coffee, tea and light meals
Luxurious and contemporary fine dining experience
Stylish retro American-style diner
Winners of the King of Curry Awards. Authentic South Indian cuisine and classic Indian food for curry lovers.
Tapas bar, Cappuccino bar and bistro serving Continental Fusion and food with a Portuguese influence.
Also known as Cafe Bahia.
Authentic and exciting Turkish food on Linthorpe Road
Italian cafe and restaurant offering quality food at great prices.
Dickens Inn is the ideal place to start your perfect night out with a relaxed beer garden or enjoy the stylish vibe of Dickens 2.
Gourmet burgers and grill, plus cocktails
The Southfield is directly opposite the Teesside University Campus. The pub has a friendly crowd with a good mix of people.
Continental bar, perfect for an afternoon rendezvous.
The Swatters Carr sells a delicious variety of food from 8am-10pm every day of the week! A must for history buffs as the walls are absolutely covered with interesting local history material.
Juice and smoothie bar
A mixture of English and Italian products freshly made for you
The best fry-up ever. Always hot, fresh and delicious.
Fresh sandwiches made in a friendly atmosphere
Middlesbrough's first mobile Pizza Slice seller serving fresh pizza on the streets of Middlesbrough!


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Revista Matemática Complutense
Revista Matemática Complutense is an international research journal publishing high quality research and survey articles in a broad spectrum of pure and applied mathematics. The fields of interests include: analysis, differential equations and applications, geometry, topology, algebra, statistics, computer sciences and astronomy. The journal is sponsored by the School of Mathematics at Complutense University in Madrid. The prestigious interdisciplinary editorial board, formed by more than 30 internationally well-recognized researchers in different areas, reflects the quality standards of our journal.
Since 2009, the journal appears listed in the Journal Citation Report.
- 6 Volumes
- 13 Issues
- 178 Articles
- 7 Open Access
- 2010 - 2015 Available between
Latest Articles
Capacities and Hausdorff measures on metric spaces
Nijjwal Karak, Pekka Koskela (September 2015)
The convenient setting for Denjoy–Carleman differentiable mappings of Beurling and Roumieu type
Andreas Kriegl, Peter W. Michor, Armin Rainer (September 2015)
An introduction to functional data analysis and a principal component approach for testing the equality of mean curves
Lajos Horváth, Gregory Rice (September 2015)
- Journal Title
- Revista Matemática Complutense
- Coverage
- Volume 23 / 2010 - Volume 28 / 2015
- Print ISSN
- 1139-1138
- Online ISSN
- 1988-2807
- Publisher
- Springer Milan
- Additional Links
Continue reading...
To view the rest of this content please follow the download PDF link above.


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USS Silversides Museum in Muskegon, Michigan will be presenting a series of films titled, Great Lakes Film Series: Storms, Shipwrecks, Rescues and History.
This series of four films will be presented on Wednesdays in July at 11:00am and then again at 2:30pm, beginning Wednesday, July 8, 2015, with Ric Mixter as creator and producer of the series of intriguing Great Lakes documentaries. He produced the outstanding video "Storm." Below is the schedule for the series:
July 8: Cutter Rescues: Video and interviews of Coast Guard Members and crews rescued during perilous storms.
July 15: Edmund Fitzgerald: History and video of the birth and then the death of this ship on November 10, 2010 on the Great Lakes.
July 22: Bombs Away: A documentary about the closure of Selfridge Air Base and the departure of the bombers is shown and narrated.
July 29: Best of the Depth: Lake Michigan
A lot of people figure that employees really don't go out of the way to do anything extra any more but Linda Falkowski who works for Consumers Energy in Muskegon, Mi has proven them wrong. Linda is a field representative for the company and as such, comes in contact with numerous customers. Because of her work ethics, Linda has been given the Gatekeeper of the Year award from the State of Michigan.
History buffs will certainly to want to check out the Pump House in Holland, MI. located directly north of the US Coast Guard Station at 2388 Ottawa Beach Rd. The Pump House will be open to the public on the third Thursday of each month from 3 – 5pm beginning June 18. The following dates will be July 16, August 20 and September 17.
Free parking is available in both the Black Lake Boardwalk's east and west parking lots at 2278 Black Lake Ave. and 2358 Ottawa Beach Rd.
Since the dedication of this building in May was so popular, the Historic Ottawa Beach Society (HOBS) and Ottawa County Parks decided to provide the public with more opportunities to view the interior of the historic Pump House building over the summer.
The art in the building will still be on the walls for guests to enjoy. The SOBS is working to set up a historical storyteller for the July through September dates.
On July 11, 2015, the Pentwater Jr. Women's Club will host the 51st Annual Pentwater Arts & Crafts Fair from 9:00am – 5:00pm. This event is to be held on the Village Green, downtown in Pentwater.
This Fair is a juried art fair. All work must be original and of the artist's own creation. Painting, ceramics, jewelry, photography, weaving, clothing, sculpture and other media will be available. People attending also have the opportunity to bid in a silent auction on art pieces donated by many of the participating artists. The proceeds generated will be used to fund a monthly mobile food pantry.
Admission is free. There will be a refreshment stand on the Village Green operated by the Pentwater Service Club.
Additional information is available by contacting the club at firstname.lastname@example.com
On Saturday, July 18, 2015, the VFW Post 4249 of Newaygo will be hosting the Michigan Knights of Columbus 4th Degree District 4, 2nd Annual Grand Rapids Home for Veterans Benefit Run.
Runners are asked to meet at 3000 Monroe Ave NE in Grand Rapids, MI at 9:00am. Ride to the VFW in Newaygo, leaving at 11:30am.
A $25 donation covers food, fun and a raffle ticket. There will also be music, prizes and refreshments.
You may call David Reese at 616-446-0726 for more information.
News Release:
On May 6, 2015, Muskegon Area Career Tech Center (MACTC) students Xavier Baron (Oakridge), Matthew Visger (Montague) and Jailon Jamison (Fruitport) traveled to Anaheim, California to compete with the brightest and most skilled computer whizzes in the nation.
The three Muskegon Area Career Tech Center (MACTC) students qualified by earning top spots at the state-level Business Professionals of America (BPA) Leadership Conference. Though the teens didn't place in the top ten this year, they enjoyed the challenge and the opportunity to visit the West Coast including a visit to Disneyland. While there, Xavier earned a Security Fundamentals and a Networking Fundamentals certification. Matthew also earned his Networking Fundamentals certification. These certifications will allow them to bypass entry-level classes and open the door to places of employment.
GRAND HAVEN, Mich., MAY 14, 2015--Since 2007, an informal group of women from St. Patrick-St. Anthony Parish has been knitting and crocheting hand made shawls, lap robes, afghans, hats, mittens, scarves and similar items to give veterans, infants, middle schoolers, hospitalized persons, those in nursing homes, parish members and community friends comfort. Now in its eighth year, the group--known as the Parish Prayer Shawl Ministry--has passed a major milestone of donating 150 items to persons in the lakeshore community and as far away as Haiti.
Hennessy's Irish Pub
Barnes & Noble Booksellers -
WZZM 13, 645 Three Mile, Grand Rapids, MI 49544
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Angry Birds Star Wars 3D 150 Pc Puzzle
Item No.1710011509
Item No.1710011509
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Star Wars Angry Birds fans will love putting together this 150 Piece Super 3D Puzzle. Based on the hit mobile phone game, this amazing 3D jigsaw features all your favourite Star Wars characters as high-flying Angry Birds and villainous pigs. The completed puzzle measures 45.7cm x 30.4cm and is suitable for kids aged three and up.
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Yasmin Rioux
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Hometown: Augsburg, Germany
Education: B.A. English; Communication from University of Dubuque, M.A. Communication from University of Dubuque
Additional M.A. work: M.A. TESOL at the Monterey Institute of International Studies in Monterey; M.A. English and Creative Writing, Southern New Hampshire University
I am also currently working on my Ph.D. in composition and TESOL at the Indiana University of Pennsylvania.
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East Lyme - There have been other times East Lyme field hockey coach Amy Lloyd thought her team was going to beat Stonington.
In 2006, Lloyd had what she thought was her best team. She even had her sister Laura at the game posing as the Viking mascot. But that day Stonington's goalie stopped a penalty stroke. And the Bears won their 16th straight Eastern Connecticut Conference title. Oh, yeah, and it was Lloyd's birthday.
"After I talked to (the newspaper), I went home and cried," Lloyd said.
And so Wednesday East Lyme beat first-place Stonington 2-1, just the second time the Vikings have won that matchup since 1990. East Lyme also won in 2005, Lloyd said.
"We knew we were kind of the underdogs," East Lyme center midfielder Paige Schneider said. "It feels great, great to get a win."
Stonington (4-3, 4-1) is still technically in first place in the ECC. Waterford (4-2, 3-1) also has one loss, while East Lyme, which stumbled early in the season as it integrated the new members of its starting lineup, is 4-2-2, 3-1-1.
East Lyme perhaps turned the corner in a Sept. 24 tie with North Branford, 2-2. Stonington lost to North Branford 1-0 on opening day.
"Rightfully so, everyone's out to beat them," said Lloyd of Stonington, which has won 20 of the last 21 ECC titles. "Sometimes it can get in the players' heads. But against North Branford we went in with a clear mind, with nothing to lose. We did that again today, we came in with a clear mind.
"Right now we have some good momentum. We're putting the ball in the cage more consistently."
Ellie Bruckner scored the game-winning goal with 5 minutes, 59 seconds remaining, assisted by Kailey MarcAurele.
The game was tied 0-0 at halftime. Stonington's Molly Crowley scored first, assisted by Chloe Slater on a penalty corner with 21:10 to go, but in all the Bears generated just 10 shots off 15 corners, bottled up by the Vikings' defense and their own execution errors.
East Lyme's Alice Navarro tied the game on an assist by Caroline Regan about four minutes later.
It was then Stonington's turn for a flurry of offense and East Lyme's Schneider, one of the Vikings' top and toughest play-makers, was forced to come out of the game with 12:39 remaining with an injury to the fingers on her left hand.
Lloyd moved Regan, a forward, to Schneider's spot in the midfield and she carried the ball in, generating the winning goal.
"That could've had an impact (losing Schneider)," Lloyd said. "Even if we were just a little off."
Regan, Schneider, Samantha Horner and Kelsey Shukis were the four players Lloyd said were most involved in breaking up Stonington's corners, something the Vikings practiced extensively leading up to the game. East Lyme "switched up" its defensive placement on corners, Lloyd said.
East Lyme's Sarah Bond made nine saves in the victory.
"We had a lot more opportunities, but we didn't handle the ball," said Stonington coach Jenna Tucchio. "We have lots of mistakes we need to address. East Lyme is a skilled team, but you need to execute the fundamental pieces of the game properly. ... You can't make any mistakes (and still win the ECC). Well, you can make one, but you can't make two."
Animal advocates monitor cruetly case of former Mystic, Niantic dog trainer
Chief Ackley's future separates the New London mayoral candidates
Electric Boat employees offer free tutoring to local students
Doctors Without Borders: 19 dead, including 3 children in Kunduz hospital bombing
Nearly a dozen arrested in prostitution, narcotics sting in Groton
Stonington High School bans grinding and other 'inappropriate dancing'
Former Hartford police officer plans to remove vessel from Crescent Beach
'Soup kitchen' could run out of operating funds by the end of the year
Newtown families' suit against gun maker sent to state court


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Keep your stuff close and your casual look tied together with this modestly sized saddle bag. The zippered fold-over flap offers the practicality of a large, secure extra pocket. The sueded tassel? That's just for fun.
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- Sueded tassel pull on zipper, pebbled faux leather
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- 5 1/4" tall x 9" wide x 1" deep
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The Lady Falcons' Class of 2016 members helped the team's three freshmen acclimate themselves to varsity high school softball
Dateline will air a sit-down interview with 27 of the women who have accused
Much like Dracula, once Ryan Murphy finds a woman he likes, he never wants to let her go. The
Following the success of The Martian, Hollywood has already released a trailer for its...
Like many other 14-year-olds, ABC Family is changing its name, and it now wants to be called Freeform, the network announced Tuesday. According to a press release, the reason behind the change is to is to better cater to "Becomers," a...
Burger King's new Halloween-themed Whopper is causing a lot of kerfuffle on the Internet, and for good reason: Apparently, the special black bun turns your poop bright green. We've reached out to good...
A cheetah cub named Kumbali and a young labrador mix named Kago have forged an unlikely friendship at Virginia's Metro Richmond Zoo.
Rihanna is defending Rachel Dolezal, saying she sees nothing wrong with the white NAACP leader
Five or six months ago, libertarians supposedly were on the march in the Republican Party. Rand Paul was one of the leaders in the fight for the GOP presidential nomination, and the push for much smaller government somehow seemed ascendant among...
October is Breast Cancer Awareness Month, and all around the nation, people are doing their part to raise awareness and support survivors. These events promote community and fun while benefitting a worthwhile cause.
Voting for the Most Popular Driver awards in NASCAR's XFINITY and Camping World Truck Series, along with NASCAR's seven touring series, began today.
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We are a family from the west, making the most of our time in the east. Loving the lush green, the four seasons, the abundant water, and the bounty of local organic and humanely-raised food. Vermont is the real deal!
I'm a modern dancer, writer, English teacher, baker, clothesline architect, novice vegetable gardener, mother of a teen and a toddler.


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College Life
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The H Sweater Is Back
If you've been on this campus for even a single mildly cold day, you've seen the H Sweater. It's either cream or crimson and it's adorned with an obnoxious, torso-sized "H," in case anyone forgot what letter this University's name starts with.
Harvard Stadium Plays Host to Weekly Stair Runs
Boston-area residents participate in weekly stair runs at the Harvard Stadium, enjoying a sense of community.
Isaac Inkeles on Social Conservatism
Isaac G. Inkeles '16 is a self described social conservative and notes that he often hesitates to share his initial opinions on social issues with more liberal students.
After Critique, College Reevaluates Hall Common Rooms
Undergraduates are critical of the hallway common rooms featured in recent House renovations at the expense of private space.
UC Leaders Draw Ire for App Funding Decision
Undergraduate Council representative criticize the Council's decision to continue funding smartphone application Ventfull.
The Elephant in the Room: Conservatives at Harvard
It is not easy being a conservative at Harvard, surrounded by a sea of blue and the tradition of a school once called the "Kremlin on the Charles." Fear of judgment and misinterpretation cause many conservative students to remain quiet on their political beliefs, or crawl into a closet with regard to their beliefs.
Chris Heller
Chris Heller, a piano-based, pop singer-songwriter, performs in the Science Center plaza at noon on Tuesday, September 29, 2015.
Senior Pictures
"I feel I have to get an official photo for my mom and to be in the yearbook." Shaquilla T.S. Harrigan '16 poses for a senior portrait by Steve E. Anderson, Senior Photographer from Prestige Portraits on Tuesday, September 29, 2015.
Admissions Office Fields Questions on Final Clubs
Dean of Admissions and Financial Aid William R. Fitzsimmons '67 said the all-male social organizations are "not consistent with the mission of the College."
Senior Portraits
"I feel I have to get an official photo for my mom and to be in the yearbook." Shaquilla T.S. Harrigan '16 poses for a senior portrait by Steve E. Anderson, Senior Photographer from Prestige Portraits on Tuesday, September 29, 2015.
Yearbook Offers Make-Up Photo Sittings for Seniors
Yearbook president Dennis Lee '16 said that he "sincerely hopes" with the additional sittings, all seniors will have the chance to be pictured in the yearbook, even if they were unable to attend their original sitting.
Popular Square Eateries' Exodus Prompts Stir Among Students
Both Al's and Clover are popular local institutions, and their forthcoming departure because of planned renovations to the Smith Campus Center, caused a stir among some students.
Pontiff's Visit Inspires Students on Trip to Philadelphia
During Pope Francis' historic visit to the United States, two Divinity School faculty members accompanied a group of more than 50 undergraduates to catch a glimpse of the leader of the Catholic Church.
Honor Council Members Adjust Schedules as Hearings Begin
Undergraduate members of the Honor Council—the student-faculty body tasked with enforcing the honor code—are adjusting their schedules as the Council hears its first slate of academic integrity cases.
Student Life Office Names Two Administrators
Loc Truong will serve as the inaugural director of the College's new Office for Equity, Diversity, and Inclusion, dedicated to supporting first generation college students, undocumented students, and foster youth.


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The F.B.I.: Season 2, Episode 28
Force of Nature (9 Apr. 1967)
Ratings: 8.5/10 from 10 users
Reviews: write review
Operating on a tip, Erskine and Rhodes arrest Charles Burnett and Allen Cole just before they were going to commit murder and kidnapping. The tipster was Burnett's former wife, for whom he ... See full summary »
Director:Jesse Hibbs
Writer:Mark Rodgers


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Josh Brent was driving as fast as 134 mph before the December crash that killed his Cowboys teammate
- Cowboys nose guard Josh Brent was caught on at least three surveillance cameras driving at a minimum of 110 mph before the crash
- Police documents show he was going as fast as 134 mph at some points
- His friend and teammate Jerry Brown Jr died when the Mercedes S600 hit a curb and flipped on December 8
- Brent is charged with intoxication manslaughter after he was found to be more than twice the legal blood-alcohol limit
Dallas Cowboys footballer Josh Brent was driving as fast as 134 mph before a drunken crash that killed his teammate Jerry Brown Jr, documents revealed today.
Brent's blood alcohol level was more than twice the legal limit when he lost control of his vehicle and it flipped on Texas Highway 114 near Irving in the early hours of December 8.
According to a police report, at least three different surveillance cameras caught the player's four-door Mercedes S600 driving at speeds of at least 110 mph on Highway 114 at 2:19 a.m.
Findings: An autopsy revealed that Dallas Cowboys player Jerry Brown, left, was sober when he died from head and neck trauma in the December crash , while his teammate Josh Brent, right, was driving while legally drunk
The report, obtained by NBC 5, says Brent could have been driving more than 20 mph faster than that at some points.
Brent and Brown had been socializing with teammates at Eddie V's in Oak Lawn at around 11:15 p.m.
From there, they went on to Beamers Nightclub, where security cameras caught them leaving at around 2 a.m.
A timeline of events reveal a 911 call was made from the in-car phone at 2:19 a.m and a passer-by called for help at 2:21 a.m. Brown was taken to Parkland Memorial Hospital where he was pronounced dead just before 3 a.m.
Brent was charged with intoxication manslaughter in December but no date has been set for his trial.
Brown's autopsy in early January revealed that the Dallas Cowboys practice squad player was sober when he was killed in the crash.
The Dallas County Medical Examiner's Office reported that Brown died of head and neck trauma when the S600 vehicle overturned.
Fatal: Police say Brent had a blood alcohol limit of 0.18 percent - more than twice the legal limit - when he crashed his Mercedes in Dallas and killed his teammate
Wreck: Tire skid marks are seen leaving the road where the car driven by Dallas Cowboys player Josh Brent crashed, killing Jerry Brown Jr
He had a dislocated neck, a severely bruised spine and a blood alcohol content of 0.056 per cent -- well below the Texas drunken driving standard of 0.08 per cent.
Police said Brent, a nose guard with the Cowboys, had a blood alcohol level twice the legal limit when he crashed in Irving, a Dallas suburb. He remains free on $100,000 bond.
The medical examiner's report stated that Brown, 25, sustained abrasions on the right and left forehead and a cut on his left eyebrow.
Neither man was wearing a seatbelt at the time of the collision, according to officials.
The autopsy report followed a grand jury in Texas that formally indicted Brent on one count of intoxication manslaughter.
Brent was charged in connection with the December 8 crash that killed his friend and Cowboys practice squad member, who was also his teammate for three seasons at the University of Illinois.
Legal woes: Brent, who is charged with intoxication manslaughter that left his close friend dead, was released from jail on $100,000 bail
Police in Irving said on December 8, Brent was speeding along Highway 114 when his vehicle struck a curb and flipped.
Intoxication manslaughter carries a maximum sentence of 20 years in prison. Brent could be eligible for probation.
Brent's blood alcohol level was 0.18, meaning a person his weight would have to drink about 20 shots over a four-hour period to reach that level.
Cowboys coach Jason Garrett said the team would continue to support Brent despite the revelation.
is a member of our family and we will continue to support him in every
way that we can and at the same time try to help other people who are in
a similar kind of situation,' he said.
A number of witnesses came forward to say that Brown was alive and screaming for help after Brent crashed the car.
Car: Brent was driving as fast as 134 mph in his four-door Mercedes-Benz S600, similar to that pictured
Stacee McWilliams, 40, from Irving, told police that when she stopped at the scene of the accident, she practically had to beg Brent to help his best friend out of the Mercedes, which was upside down and on fire.
She said: 'I jumped out and ran out toward the wreck. I yelled to Brent, "Are you OK? Are you OK?" And he told me he was fine as he was pacing back and forth.
'I was relieved because I thought, it's just a single-car accident, nobody's hurt. And then fire sparked up from the car and got bigger and I started to hear screams.
Brent's attorney, George Milner, called McWilliams' account 'absolutely false'.
'What I've been told is that Jerry Brown was never conscious after the accident,' Milner said. 'This woman, while present in the area of the accident, could not have seen or heard anything she said about Josh.'
The two close friends started their night at Eddie V's restaurant in Oak Lawn before moving on to a night club in northwest Dallas, CBS DFW reported.
The Texas Alcoholic Beverage Commission has launched an investigation to determine whether the two football players were over-served, and if so, by whom.
Memorial: Brown's Dallas Cowboys jersey is carried into the Oak Cliff Bible Fellowship education center
Sadness: Brown's relatives embraced ahead of a memorial service for the 25-year-old player
Eddie V's released a statement, saying in part that the establishment takes 'the responsibility that comes with serving alcohol very seriously.'
It was not Brent's first run-in with the law - or intoxication charge.
In February 2009, Brent was arrested in February 2009 near the Illinois campus for driving under the influence, driving on a suspended license and speeding, according to Champaign County, Illinois.
In June 2009, he pleaded guilty to DUI and was sentenced to 60 days in jail, two years of probation, 200 hours of community service and a fine of about $2,000.
As part of his plea deal, prosecutors dropped one count of aggravated DUI/no valid driver's license. Brent successfully completed his probation in July 2011, court records show.
The Cowboys had signed Brown to their practice squad on October 24, but he hasn't been on the active roster. He was released from the Colts' practice squad on October 20. Brown played in one game for the Colts, a loss to the New York Jets on October 14.
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text: There's something unique about the dining experience with Oren. The food, made with the freshest of ingredients and supreme skill, without doubt plays the starring role. Gert is probably one of the best chefs I've encountered in Wales, he flavours his food with a deft, balanced hand. However... read more


text:
We had company this weekend and I made a German Pancake for Sunday brunch. I haven't made one in many years. Now I can't remember why it fell off my repertoire.
Every pancake is different. It's baked in the oven in a cast iron skillet. Mine didn't rise like it normally does - but it still tasted delicious. It was especially wonderful with butter, natural syrup and Jane's Good Food - Spiced Vanilla Pears. Yum!
Recipe notes: I did not use any of the fruit toppings recommended in the German Pancake recipe. I simply made the basic pancake and served it with my own homemade applesauce and the aforementioned Spiced Vanilla Pears. Also, I saw no reason to add confectioner's sugar on top of the pancake.


text:
Jun 02, 2005
NICOSIA [MENL] -- An Iranian energy firm reports an increase in crude oilproduction capacity.
May 23, 2005
LONDON [MENL] -- OPEC has again raised the prospect of a slash in oilexports.
May 18, 2005
WASHINGTON [MENL] -- Saudi Arabia would become a more vital oil supplier tothe United States and the West.