Unfortunately, after publication of this article , an error was discovered in Fig. 12 (Fig. 1 here) that was introduced during the Production process. The corrected figure can be seen below and the original article has also been updated to reflect this change.
The online version of the original article can be found under doi:10.1186/s13024-016-0078-3.
The online version of the original article can be found at http://dx.doi.org/10.1186/s13024-016-0078-3.