*1: With Tritomaria exsectiformis on peat on rocky slope, Carn Galver, Penwith, Land's End Peninsula, 1987, JAP (BBSUK) (Long 1988: 32).
*2: S. of Temple, Bodmin Moor, 1964, JAP(BBSUK) (Paton 1969a: 704).
Habitat notes from Cornwall are as follows.Lower Bostraze: on clayey soil of slope above disused china clay quarry, open or partly shaded (with Nardia scalaris). Stannon: on gritty soil of near-vertical bank amongst china clay spoil, almost unshaded (with Lophozia incisa Nardia scalaris). Rusey: soil high on heathy slope of sea-cliff.
Only seen with foliar gemmae.