In the 16th century, Babur, a Timurid descendant of Timur and Genghis Khan from Fergana Valley (modern-day Uzbekistan), swept across the Khyber Pass and founded the Mughal Empire, covering India, along with modern-day Afghanistan, Pakistan and Bangladesh The Mughals were descended from Persianised Central Asian Turks (with significant Mongol admixture). In the Mughal era, Uttar Pradesh became the heartland of the empire. Mughal emperors Babur and Humayun ruled from Agra.