The ecoregion forms a long band, about 2,100 km long and 600 km wide, from Ukraine in the west to western edge of Siberia at the Ural Mountains in the east. Most of the terrain is rolling hills and some plains. Average elevations range from 150 meters (above sea level) in the lowlands to 250 meters in the hills, with some isolated ranges of low mountains on the edges of the area. The ecoregion can be divided into sub-provinces from west to east: Dnieper Upland, Dnieper Lowland, Central Russian Upland, Oka-Don Plain, Volga Upland, and the Trans-Volga. Moisture diminishes as the ecoregion stretches to the east, as does the forest fragmentation.