A number of lakes formed or expanded in the Sahara. The largest of which was Lake Chad which increased to at least ten times its present-day size to form Lake Megachad. This enlarged Lake Chad reached dimensions of in north-south and east-west direction respectively, covering the Bodélé Depression and perhaps as much as 8% of the present-day Sahara desert. It influenced the climate itself; for example rainfall would have been reduced at the centre of the lake and increased at its margins. Lake Chad was possibly fed from the north by rivers draining the Hoggar (Taffassasset drainage) and Tibesti Mountains and from the south by the Chari-Logone and Komadugu Rivers. The Chari River was the main tributary while the rivers draining the Tibesti formed alluvial fans/the Angamma river delta at their entry into northern Lake Chad. Skeletons of elephants, hippos and hominins have been found in the Angamma delta, which is the dominant shoreline feature of northern Lake Chad. The lake overflowed into the Niger River during highstand through the Mayo Kebbi and the Benue River, eventually reaching the Gulf of Guinea. Older dune systems were submerged by Lake Chad.