Wildlife crossings have also been important for protecting biodiversity in several areas of southern California. In San Bernardino County, biologists have erected fences along State Route 58 to complement underpasses (culverts) that are being used by the threatened desert tortoise. Tortoise deaths on the highway declined by 93% during the first four years after the introduction of the fences, proving that even makeshift wildlife crossings (storm-drainage culverts in this case) have the ability to increase highway permeability and protect sensitive species . Additionally, studies by and report that underpasses in Orange, Riverside, and Los Angeles Counties have drawn significant use from a variety of species including bobcats, coyotes, gray fox, mule deer, and long-tailed weasels. These results could be extremely important for wildlife conservation efforts in the region's Puente Hills and Chino Hills links, which have been increasingly fragmented by road construction Los Angeles County's first wildlife-purpose built underpass is at Harbor Boulevard.[ It was built in partnership between Los Angeles County, California State Parks and the Puente Hills Habitat Preservation Authority.