Juglans californica is generally found in the southern California Coast Ranges, Transverse Ranges, and Peninsular Ranges, and the Central Valley. It grows as part of mixed woodlands, and also on slopes and in valleys wherever conditions are favorable. It is threatened by development and overgrazing. Some native stands remain in urban Los Angeles in the Santa Monica Mountains and Hollywood Hills. J. californica grows in riparian woodlands, either in single species stands or mixed with California's oaks (Quercus spp.) and cottonwoods (Populus fremontii).