A survey of bird populations in Saint Croix observed that Hugo's aftermath may have stressed birds more than the hurricane's immediate meteorological forces. Frugivorous, nectarivorous, and seminivorous bird populations declined most among avian diet groups as a result of vegetation loss. The bridled quail-dove (Geotrygon mystacea) was driven out of its traditional habitats on Saint Croix. Declines in the populations of certain bird species were also noted in Saint John. The destruction of habitats forced the relocation of some avian species such as the pearly-eyed thrasher (Margarops fuscatus) and northern waterthrush (Seiurus noveboracensis). The populations or habitats of three endangered Puerto Rican birds were affected by Hugo: the Puerto Rican amazon (Amazona vittata), the Yellow-shouldered blackbird (Agelaius xanthomus), and the Puerto Rican plain pigeon (Columba inornata wetmorei). El Yunque National Forest lost 15 percent of its trees, valued at $5.2 million. However, the increased exposure to sunlight following the loss of tree canopies led to increased diversity of plant species. In Montserrat, the endemic Montserrat oriole (Icterus oberi) was driven out of the South Soufriere Hills after losing much of its habitat.