 Pawley (2006) also found lysorophians to be the closest relatives of lissamphibians, but found aïstopods and adelogyrinids rather than microsaurs to be the second most closely related groups.Marjanović (2010) found holospondyls to be the most closely related group to lissamphibians, followed by lysorophians.Under this hypothesis, lepospondyls would be crown tetrapods and temnospondyls would be stem tetrapods.