 Colonies were founded at Akrai (664 BC), Kasmenai (643 BC), Akrillai (7th century BC), Helorus (7th century BC) and Kamarina (598 BC).The descendants of the first colonists, called "Gamoroi", held power until they were expelled by the "Killichiroi", the lower class of the city.The former, however, returned to power in 485 BC, thanks to the help of Gelo, ruler of Gela.